Glacial and post-glacial adaptations of hunter-gatherers: Investigating
the late Upper Paleolithic and Mesolithic subsistence strategies in the
southern steppe of Eastern Europe
Keiko Kitagawa
a, *
, Marie-Anne Julien
a, b
, Oleksandra Krotova
c
,
Alexander A. Bessudnov
d
, Mikail V. Sablin
e
, Dmytro Kiosak
f
, Natalia Leonova
g
,
Boris Plohenko
g
, Maryl
ene Patou-Mathis
a
a
Unit e Histoire Naturelle de l’Homme Pr ehistorique (UMR 7194), Sorbonne Universit es, Mus eum National d’Histoire Naturelle, CNRS, 1 rue Ren e Panhard,
75013, Paris, France
b
Centre for the Archaeology of Human Origins, Archaeology Department, University of Southampton, Avenue Campus, Southampton, SO17 1BF, United
Kingdom
c
National Ukrainian Academy of Science, Institute of Archaeology,12, Geroiv Stalingrada ave., 04210, Kyiv e 210, Ukraine
d
Institute for the History of Material Culture, Russian Academy of Sciences, Dvortsovaya Nab.18, 191186, Saint Petersburg, Russia
e
Zoological Institute, Russian Academy of Sciences, Universitetskaya Nab. 1, 199034, Saint Petersburg, Russia
f
Department of Archaeology and Ethnology of Ukraine, I.I. Mechnikov State University, Dvoryanskaya 2, UA-65082, Odessa, Ukraine
g
Lomonosov Moscow State University, Faculty of History, Department of Archaeology, GSP-1, Lomonosovsky Prospekt, 27-4, Moscow,119991, Russia
article info
Article history:
Received 20 June 2016
Accepted 8 January 2017
Available online 18 May 2017
Keywords:
Late Upper Paleolithic
Mesolithic
Subsistence practices
Steppe
Hunter-gatherers
abstract
Diverse landscapes and ecosystems stretching across Europe led to diverse hunter-gatherer cultural
records during the Upper Paleolithic and Mesolithic. In response to abrupt climatic forcing, starting
around the Late Glacial Maximum and followed by climatic events such as the BøllingeAllerød and the
Younger Dryas in the Terminal Pleistocene, archaeological data pertaining to cultural and behavioral
shifts of hunter-gatherers continue to be explored on a regional and pan-regional scale. Here we present
an initial summary, which includes new and published data on faunal analyses from multiple open air
sites that span the Late Pleistocene to the Holocene, dated between the Late Upper Paleolithic and
Mesolithic (20,000e6000 uncal
14
C BP) in the southern steppe of Eastern Europe. For this area, this is the
first study to compile the cultural and faunal data with geo-referenced localization and radiometric dates
of the archaeological sites. Taken together, faunal assemblages from the Epigravettian are characterized
by low diversity and are often dominated by one species of large game, including bison and equids,
whereas the Mesolithic diet is characterized by higher inter-site variability subsisting on large ungulate
and greater emphasis on freshwater resources. This study contributes to the general knowledge con-
cerning the last phases in the evolution of the Eurasian hunter-gatherers.
© 2017 Published by Elsevier Ltd.
1. Introduction
The past adaptation of Paleolithic and Mesolithic hunter-
gatherers in Europe is a result of their interaction with the
diverse landscapes and ecosystems. Unlike other cultural
transitions where the relationship between climatic forcing and
human behavior remains to be demonstrated, the shift from the
Late Upper Paleolithic and Mesolithic has been linked to and
framed in terms of abrupt climatic changes occurring between the
Late Glacial and the Holocene. In response to abrupt climatic forc-
ing starting around the Late Glacial Maximum and followed by
climatic events such as the BøllingeAllerød and the Younger Dryas
in the Terminal Pleistocene, archaeological data pertaining to cul-
tural and behavioral shifts of hunter-gatherers have been explored
on a regional and pan-regional scale (Cromb e et al., 2011; Donahue
and Lovis, 2006; Huntley et al., 2013; Lovis et al., 2006). Within the
* Corresponding author.
E-mail addresses: kkitagawa@gmail.com, keiko.kitagawa@mnhn.fr
(K. Kitagawa), M.Julien@soton.ac.uk (M.-A. Julien), okrotova@ukr.net (O. Krotova),
bessudnov_a22@mail.ru (A.A. Bessudnov), msablin@yandex.ru (M.V. Sablin),
dkiosak@ukr.net (D. Kiosak), nbleonova@gmail.com (N. Leonova),
koschmarik2008@yandex.ru (B. Plohenko), patmath@mnhn.fr (M. Patou-Mathis).
Contents lists available at ScienceDirect
Quaternary International
journal homepage: www.elsevier.com/locate/quaint
http://dx.doi.org/10.1016/j.quaint.2017.01.005
1040-6182/© 2017 Published by Elsevier Ltd.
Quaternary International 465 (2018) 192e209