American Journal of Applied Sciences 6 (6): 1076-1085, 2009
ISSN 1546-9239
© 2009 Science Publications
Corresponding Author: Norma Almaraz-Abarca, Interdisciplinary Research Centre for Integral Regional Development-Durango,
National Polytechnique Institute (CIIDIR-IPN-Dgo.), Sigma s/n, Durango, Dgo., Mexico, 34220
Tel/Fax: (52 618) 8142091
1076
Profiling of Phenolic Compounds of Somatic and Reproductive
Tissues of Agave Durangensis Gentry (Agavaceae)
Norma Almaraz-Abarca, Elí Amanda Delgado-Alvarado, Vicente Hernández-Vargas,
Margarita Ortega-Chávez, Gildardo Orea-Lara, Armando Cifuentes-Díaz de León,
José Antonio Ávila-Reyes and Raúl Muñiz-Martínez
Interdisciplinary Research Centre for Integral Regional Development-Durango,
National Polytechnique Institute (CIIDIR-IPN-Dgo.), Sigma s/n, Durango, Dgo., Mexico, 34220
Abstract: Problem statement: In Durango, Mexico, mescal is elaborated from wild plants of Agave
durangensis. This species shows a high morphological variability within and among populations, what
makes its taxonomic delimitation a hard task. Approach: In this study the pollen and foliar phenolic
compositions of Agave durangensis were analyzed by HPLC/DAD with the aim of determining the
significance of phenol profiles to delimit this taxon. Results: The foliar phenol compositions were
evaluated within and among two populations and between juvenile and adult plants. Agave asperrima
Jacobi, Dasylirion sp. and juvenile samples of A. shrevei Gentry subsp. shrevei, A. shrevei Gentry
subsp. matapensis Gentry and A. wocomahi Gentry, were also analyzed to stand comparisons with. The
results from this study indicated that pollen and foliar tissues of Agave durangneisis were rich in
kaempferol glycoside derivatives (13 and 23 different compounds can be present, respectively).
Principal coordinates analysis (PCO), based on foliar profiles of adults, indicated the presence of
several chemotypes within the Type locality of Agave durangensis and revealed chemical differences
between the both analyzed populations. Conclusion/Recommendations: Chemical and morphological
differences and biogeographical evidence suggest the recognition of two different taxonomic entities in
this morphological variable group.
Key words: Pollen flavonoids, foliar flavonoid profiles, Agave phenolic variability
INTRODUCTION
Agave is the biggest genus of the family
Agavaceae, with around 166 species, from which
125 grow in Mexico
[1,2]
. Relevant ethnobotanic
relationships have been established between the
elements of this genus and the ancient and present
cultures of Mexico
[3,4]
. Several authors have described
the use of Agave as source of fibers, food and
beverages
[5]
. In addition, Agave is used as natural
fences to avoid the soil erosion and as cattle food
[6]
.
At the present, the relevance of Agave has
increased meanly because of the increased demand of
alcoholic beverages like tequila and mescal
[7]
and the
research on potential sources of prebiotics
[8]
. In all the
cases, the authentification of the species of Agave to be
used is an important requirement in the quality control
of the manufacturing processes. In Mexico, with
exception of Agave tequilana Weber var. azul, species
like Agave durangensis, which support local industries
of alcoholic beverages and which with just begin an
industrialization process, techniques are needed to
guarantee the botanical origin of plants, according to
the respective origin denomination statement.
Agave durangensis Gentry is one of the 24 species
of genus Agave occurring in Durango, Mexico
[9]
. This
species belongs to group Ditepalae of subfamily
Agavoideae, proposed by Gentry
[3]
and can be found
from Southern Durango to Northern Zacatecas
[3,10]
.
In two localities of Durango (“Sierra of Registro”,
the type locality and “Mezquital”), populations of
Agave, traditionally called “agave mezcalero” or ”agave
cenizo”, presumably A. durangensis, maintains a
thriving mescal industry. Actually, studies have not
been done to confirm that in all the cases the raw
material to elaborate mescal is composed of that
species
[10]
. In Durango, the manufacturing of mescal is
mainly based on the gathering of agaves from their
natural populations
[7]
. Recently, producers have been
interested in the establishment of plantations of