Journal of Plant Diseases and Protection, 113 (4), S. 188–189, 2006, ISSN 1861-3829. © Eugen Ulmer KG, Stuttgart
J.Plant Dis.Protect. 4/2006
Evaluation of Solanum rootstock accessions for control of root-knot nematodes and
tobamoviruses
Bewertung von Solanum-Veredelungsunterlagen hinsichtlich ihrer Resistenz gegenüber Wurzelgallennematoden und
Tobamoviren
E.A. Tzortzakakis
1*
, F.A. Bletsos
2
& A.D. Avgelis
3
1
N.AG.RE.F, Plant Protection Institute, Nematology Lab, PO BOX 2228, GR-71003, Heraklion, Crete, Greece
2
N.AG.RE.F, Agricultural Research Center of Macedonia and Thrace, Department of Vegetables, PO BOX 60458, GR-57001, Thermi-Thessaloniki, Greece
3
N.AG.RE.F, Plant Virus Lab, GR-71003, Heraklion, Crete, Greece
*
Corresponding author, e-mail etzortza@her.forthnet.gr
Received 16 March 2006; accepted 13 April 2006
Summary
Twenty-two Solanum accessions used as rootstocks were
evaluated for their resistance towards the root-knot nema-
todes Meloidogyne javanica and M. incognita and three tobam-
oviruses (TMV, ToMV and PMMV). Five out of seven S. gilo
accessions, namely RNL 300/328, RNL 304/337, RNL
308/347, RNL 297/323 and RNL 307/346, were resistant or
immune against all viruses while others within S. aethiopicum
group kumba, S. linnaeanum, S. macrocarpon and S. dasyphyl-
lum were susceptible. Furthermore, variability of plants with-
in the same accession being either susceptible or resistant/
immune to viruses was recorded in others. Both nematode
species produced galls and egg masses on all tested plants.
Key words: Meloidogyne javanica, Meloidogyne incognita,
pepper mild mottle virus (PMMV), tobacco mosaic virus
(TMV), tomato mosaic virus (ToMV)
Zusammenfassung
Zweiundzwanzig Veredelungsunterlagen der Gattung Sola-
num wurden hinsichtlich ihrer Resistenz gegenüber drei Toba-
moviren (TMV, ToMV and PMMV) und den Wurzelgallenne-
matoden Meloidogyne javanica und M. incognita bewertet. Die
fünf (von sieben untersuchten) S.-gilo-Unterlagen RNL 300/
328, RNL 304/337, RNL 308/347, RNL 297/323 und RNL 307/
346 erwiesen sich als resistent oder immun gegenüber allen
Viren, während andere Unterlagen der Arten S. linnaeanum,
S. macrocarpon, S. dasyphyllum und der S.-aethiopicum-Gruppe
kumba anfällig waren. Andere Unterlagen derselben Akzession
variierten darüber hinaus in ihrer Anfälligkeit oder Resistenz/
Immunität gegenüber diesen Viren. Beide Nematodenarten
bildeten Gallen und Eimassen an allen überprüften Pflanzen.
Stichwörter: Meloidogyne javanica, Meloidogyne incognita,
Mildes Paprikascheckungsvirus (PMMV), Tabakmosaikvirus
(TMV), Tomatenmosaikvirus (ToMV)
Grafting vegetables on resistant rootstocks is an effective
method to control soil-borne pathogens in areas where they
are intensively grown. Experiments with grafted eggplants
showed promising results for control of fungal pathogens
(IOANNOU 2001; BLETSOS et al. 2003; BLETSOS 2006) and
root-knot nematodes (IOANNOU 2001) under greenhouse
conditions in Greece and Cyprus.
According to our knowledge, information on the response
of Solanum rootstocks is limited for root-knot nematodes and
absent for tobamoviruses. Several common cultivated hybrids
of tomato and eggplant in Crete are resistant to tobamoviruses
but not to root-knot nematodes. For successful grafting and
economic plant production, the rootstocks should be resistant
to both root-knot nematodes and tobamoviruses. The latter is
important, as an infection by soil-borne viruses will cause a
hypersensitive reaction in the rootstock with detrimental
effects on the grafted virus–resistant plant. Therefore, Solanum
accessions (regenerated by F. A. Bletsos, E. U. EGGNET Project)
were evaluated against tobacco mosaic virus (TMV), tomato
mosaic virus (ToMV) and pepper mild mottle virus (PMMV).
Furthermore, the ability for reproduction of one Cretan popula-
tion each of M. javanica and M. incognita was tested. The aim
of the work described here is to assess the potential use of
these Solanum accessions as rootstocks for grafting tomato and
eggplant, as the above mentioned viruses and root-knot nema-
todes are widespread in Crete causing severe economic losses.
Nematode resistance was evaluated as follows. Egg masses
of each nematode species were picked from roots of galled
tomatoes, placed on 90 Pm mesh sieves sited over water in
shallow trays and incubated at 25°C (HUSSEY and BARKER
1973). Hatched second-stage juveniles (J
2
) collected within
four days were used as inoculum and pipetted in 300 ml pots
with seedlings at the 2-4 true leaf stage at a rate of c. 300 J
2
per plant. The plants were grown for 6 weeks in a climate
room with 16 h photoperiod and 24-26°C soil temperature.
Plants were watered and fertlized as required. The number of
galls and egg masses was assessed in 4-5 replicates under a
stereoscope. In each experiment 3-4 accessions were tested
together, with controls being either the susceptible tomato cv.
‘Ace’ or the eggplant cv. ‘Tsakoniki’. Means were compared
with their respective controls by t-test at P d 0.05 and results
were presented as percentage of galls and egg masses on each
Solanum accession compared with the control.
For virus transmission, 10-15 seedlings from each accession
at the third leaf stage were mechanically inoculated at the
cotyledons and the first true leaf with a purified virus suspen-
sion (0.1 mg/ml virus nucleoprotein). After one month the
presence of virus symptoms was detected macroscopically
and confirmed with both serological (DAS-ELISA) and biolog-
ical tests. For biological detection, mechanical inoculation
was done on test plants which were Nicotiana glutinosa for
TMV and ToMV and N. rustica for PMMV. The S. melongena cv.
‘Tsakoniki’ served as control for TMV and ToMV while Capsi-
cum annuum cv. ‘Lamuyo’ for PMMV.
Both nematode species formed galls in all tested plants. The
number of galls on tomato and eggplant ranged from 18-45
per root for M. javanica and 17-53 for M. incognita. The
respective number of egg masses ranged from 9-30 for
M. javanica and 10-38 for M. incognita. In some Solanum
accessions the number of galls and egg masses were signifi-
cantly lower than in the controls while for others they were
similar or significantly higher (Table 1).
Short Communication