Fresenius J Anal Chem (1995) 352:43-48 Fresenius' Journal of
© Springer-Verlag 1995
Limitations in the use of horseradish peroxidase
as an enyzme probe in the development
of a homogeneous immunoassay for aflatoxin B1
D. Ramana l, A. K. Capoor 1, R. B. Sashidhar 1, Ramesh V. Bhat 2
1 Department of Biochemistry, University College of Science, Osmania University, Hyderabad-500 007, India
2Food and Drug Toxicology Research Centre, National Institute of Nutrition, Hyderabad-500 007, India
Received: 25 July 1994/Accepted: 24 November 1994
Abstract. Horseradish peroxidase (HRPO) was used as a
probe to quantitate aflatoxin B 1 by a homogeneous im-
munoassay. The conjugation of AFB~ to HRPO resulted
in 54% loss of enyzme activity. In the presence of AFB1
specific antibodies, the HRPO-AFB1 conjugate showed
reversal of its lost enzyme activity by 12%. This positive
modulatory effect of antibody on the enzyme activity was
used as an analytical tool to quantitate AFB~. The homo-
geneous assay carried out with free AFB~ and HRPO-
AFBI conjugate in the presence of antibodies indicated
poor linearity as compared to the heterogeneous assay. It
was observed that the number of HRPO-lysine residues
involved in AFB~ conjugation were 6-8. The low level of
modulation of enzyme activity by antibody with respect to
HRPO-AFB 1 conjugate, could possibly be attributed to
the limited number of lysine residues in the HRPO mole-
cule and its proximity to the active site of the enzyme.
Thus, HRPO was found to be limiting as an enzyme with
respect to the homogeneous enzyme immunoassay for
AFB1 analysis. The antibodies raised were specific for
AFB1, and showed excellent linearity even at high dilu-
tion for the detection of AFB1 by ELISA indicating that
antibodies per se were not the limiting factor.
Introduction
Among the fungal toxins, aflatoxins produced by the As-
pergillusflavus and Aspergillus parasiticus group of fungi
have received world wide attention, due to their deleteri-
ous effects on human and animal health and their impor-
tance in international trade. The toxicological properties
of aflatoxins include carcinogenic, mutagenic, teratogenic
and immunosuppressive effects [1-4]. Earlier, various an-
alytical methods have been developed for screening and
detection of aflatoxins in food, feed and biological fluids.
These methods include physico-chemical methods such as
Correspondence to. R. B. Sashidhar
TLC, HPLC, GLC and HPTLC and immunoanalytical
methods like RIA and ELISA for AFB~ [5-9].
In the recent past, methods based on ELISA have also
been collaboratively evaluated for aflatoxin (AFB~) analy-
sis [10]. So far, all the enzyme immunoassays developed
for AFB1 quantitation have been based on solid-phase im-
munoassay systems. However, no enzyme-mediated im-
munoassay for AFB~ based on a separation-free system
has been reported. The objective of the present communi-
cation was to develop a homogeneous (separation-free)
immunoassay for the detection of AFB~ based on an en-
zyme-modulator mediated assay using AFB1 specific poly-
clonal antibodies, with horseradish peroxidase (HRPO) as
the enzyme probe.
Materials and methods
Reagents
Aflatoxin B~ and carboxymethoxylamine hemichloride
were obtained from Aldrich Chemical Co., USA. Bovine
serum albumin (fatty acid free), poly-D-lysine, horseradish
peroxidase type VI (EC 1.11.1.7.), tributylamine, isobutyl
chloroformate, thimerosal, 1-ethyl-3-(3-dimethyl-amino-
propyl) carbodiimide hydrochloride, o-phenylene diamine
and anti-rabbit IgG (raised in goat) conjugated to alkaline
phosphatase were purchased from Sigma Chemical Co.,
USA. N-hydroxy succinimide and dicyclohexyl carbodi-
imide were obtained from Spectrochem, India. Freund's
complete and incomplete adjuvants were obtained from
Difco Laboratories, USA. Polystyrene microtiter ELISA
plates were procured from Nunc., Denmark. All other
chemicals and buffer salts were of analytical grade.
Preparation of antigen." BSA-AFB~ conjugate
Aflatoxin B1 was activated by the addition of a carboxyl
group to the molecule by the methods of Langone and Van
Vunakis [11]. Briefly, 5 mg (16 ~tmol) of AFB~ and 7.6