Introduction
Collembola (springtails) are not a well-known group because of
their small size and cryptic habits. They are, however, extremely
abundant and widespread, being the most numerous and diverse
arthropods in soils and leaf litter after mites and, in some
situations, outnumbering them. Taxonomically, the Collembola
are a class of arthropods that are morphologically similar to
insects in gross body structure, but different in the lack of a hard
exoskeleton and wings, in the presence of internal mouthparts
and in possessing, primitively, only a few simple eyes on each
side of the head and a jumping organ located ventrally. These
differences make their collection and treatment taxonomically
often quite different from insects.
As detritivores, Collembola play an important role in the
decomposition of dead plant material and in nutrient cycling.
They are generally considered to be beneficial organisms as few
species feed on live plants (Hopkin 1997). Average densities in
soil are between 5000 and 30000 per m
2
, rarely reaching close to
100000 per m
2
, and there could be up to 50 species in each
square metre (Table 1) (Greenslade 1993). As well as being
common under stones, in soil, humus and the leaf litter layer,
these animals may also be found above the ground on grasses,
under bark on trees, on flowers and leaves in tree canopies and on
shrubs and in rotting timber. They occur in extreme habitats such
as caves and on the seashore. As these animals are so widespread
and abundant, occurring from polar regions to arid and tropical
regions, their influence in ecosystem functioning is considerable.
The Australian fauna is not well known. About 400 species
are described; a further ~1000 are known but not described. The
fauna probably consists of more than 2000 species (Greenslade
1994a). Australian endemism is high at ~80% and local
endemism is significant in some groups. In addition to the
indigenous fauna, there is a substantial group of introduced,
exotic organisms that were probably imported with soil, plants
and stock, mainly from Europe, during the early days of
European settlement.
Any bioindicator that can easily and rapidly measure the
effects of impacts may contribute to management of the
ecosystem to improve sustainability. Impacts are those factors
that are deleterious to the ecological sustainability of
ecosystems. In this paper, I first highlight the characteristics of
springtails that have made them useful as indicators for the
effects of a range of natural and human induced impacts to the
environment, second, critically describe methods of collection,
including a protocol for an intensive survey of the fauna, and
third, describe field work in Australia where they have proved of
value in measuring the effects of environmental impacts.
Collembola as an indicator group
To be useful in measuring environmental change, an indicator
must provide measurable and repeatable information in
response to whatever question is being posed about the
environment more rapidly than would be obtained by a
superficial visual inspection. Ideally, it should also be able to
predict trends, by responding linearly to the impact being
studied. Over the last 20 years or so, authors of several papers
have listed the general characteristics of an ecological indicator
(Majer 1983; Greenslade and Greenslade 1984; Andersen 1999;
Büchs 2003b). Andersen (1999) focuses on the problems of
selection and validation, and while Büchs’ (2003b) treatment is
thorough, this author gives no examples of Collembola and
concentrates on European examples of other invertebrates. All
authors have stressed that the main traits required in an indicator
taxon are sensitivity, responsiveness, ubiquity, being
functionally important in ecosystems and representative, that is
being able to act as a surrogate for other taxa, as well being easy
to sample, identify and analyse. A range of invertebrates has
been proposed as indicators for Australia (Neville and Yen 2007;
New 2007). The Collembola are one such group, together with
ants, mites, herbivorous bugs, spiders and carabids. As might be
expected, no single group is ideal for all situations, but the
Collembola have proven to be a robust indicator in a range of
ecosystems in Australia if certain parameters such as the
appropriate collecting method and level of identification, are
applied correctly. Collembola are present in all ecosystems, are
0816-1089/07/040424 10.1071/EA05264 © CSIRO 2007
Penelope Greenslade
Department of Botany and Zoology, Australian National University, Canberra, ACT 0200, Australia.
Email: penny.greenslade@anu.edu.au
Abstract. Collembola have been used to assess the ecological status of a range of ecosystems in Australia over the last
35 years. The traits that Collembola possess that make them valuable as ecological indicators are listed. Examples of
situations where they have been used to measure impacts in mainly agricultural landscapes are described and the
contributions the results have made to management for ecological sustainability are noted. Densities of exotic and native
species in a variety of ecosystems are provided and a list of genera with exotic species in Australia is given. A protocol for
comprehensively surveying Collembola is described and a critique of different methods given.
The potential of Collembola to act as indicators
of landscape stress in Australia
Australian Journal of Experimental Agriculture, 2007, 47, 424–434 www.publish.csiro.au/journals/ajea
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