Formation of Volatiles in the Lipoxygenase Pathway as Affected by Fruit Type
and Temperature
Yafei Han and Sheryl Barringer
*
Department of Food Science and Technology, The Ohio State University, 2015 Fyffe Road, Columbus, OH 43210, USA.
*
Correspondence author: Sheryl Barringer, Department of Food Science and Technology, The Ohio State University, 2015 Fyffe Road, Columbus, OH 43210, USA. Tel:
614-688-3642; E-mail: barringer.11@osu.edu
Received date: Sep 16, 2015; Accepted date: Nov 2, 2015; Published date: Nov 9, 2015
Copyright: © 2015 Han Y, et al. This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use,
distribution, and reproduction in any medium, provided the original author and source are credited.
Abstract
Objective: The objective of this study is to study the effect of temperature, lipoxygenase activity, substrate and
fruit type on formation of volatiles in the LOX pathway.
Methods: Volatiles in the headspace of tomato, bell pepper, tomatillo, strawberry, apple and pear were measured
during and after fruit tissue disruption in real time using selected ion flow tube mass spectrometry (SIFT-MS) at 4,
25, 37 and 45°C for 60 min, and were compared to lipoxygenase (LOX) activity and fatty acid concentration.
Results: In tomatoes, bell peppers and tomatillos, (Z)-3-hexenal, (E)-2-hexenal and hexanal increased rapidly
after blending, peaked and declined, while in strawberry, apple and pear they sometimes peaked and sometimes
remained constant over the 60 min. In tomato and bell pepper, the peak levels of (Z)-3-hexenal, (E)-2-hexenal and
hexanal in the LOX pathway had good correlations to LOX activity, which was greatest at 25°C. In contrast, tomatillo,
strawberry, apple and pear had lower LOX activity, their 6C aldehydes did not show strong correlation with LOX
activity, and the volatiles increased with increasing temperature to 45°C. The concentrations of hexen-1-ol and 1-
hexanol did not correlate to LOX activity for any of the fruits. In tomato, bell pepper and tomatillo, there was no
significant correlation between the ratio of linoleic acid:linolenic and the ratio of hexanal:hexenal. In strawberry,
apple and pear, there were good correlations between linolenic acid content and concentrations of (E)-2-hexenal,
hexenal and hexen-1-ol.
Conclusion: Thus the effect of enzyme activity and substrate composition on formation of volatiles in the
lipoxygenase pathway remains complicated and difficult to predict.
Keywords: Lipoxygenase; Temperature; Tomato; Bell pepper;
Tomatillo; Strawberry; Apple; Pear; Selected ion fow tube mass
spectrometry
Introduction
As aroma is one of the most important characteristics for fruit
quality, volatile compounds are likely to have a huge efect on the
perception and liking of fruit products by consumers. In fruits,
important aroma compounds are derived from fatty acids, amino
acids, phenols and terpenoids [1]. Tus the efect of enzyme activity
and substrate composition on formation of volatiles in the
lipoxygenase pathway remains complicated and difcult to predict.
Among these compounds, volatiles generated from fatty acids via the
lipoxygenase (LOX) pathway are responsible for the “green” fresh note,
which is desirable in many fruit products [2]. Te LOX pathway is an
enzymatic pathway for lipid oxidation. Te main products of the LOX
pathway are C6 aldehydes and alcohols generated through sequential
enzymatic reactions. Te major enzymes in this pathway include
lipoxygenase, hydroperoxide lyase (HPL), alcohol dehydrogenase
(ADH) and (Z)-3/(E)-2 isomerase [3]. Linoleic and linolenic acids are
the substrates that are converted to volatiles such as hexanal, (Z)-3-
hexenal, (E)-2-hexenal, 1-hexanol, hexen-1-ol, (E)-2-octenal, (E)-2-
heptenal, (E)-2-pentenal 1-penten-3-one, hexanoic acid, (Z)-3-hexenyl
acetate and hexyl acetate.
Temperature afects the volatile concentrations in the headspace,
due to two main factors: diferent partition coefcients at diferent
temperatures, and diferent enzymatic activities at diferent
temperatures. Te partition coefcient is the ratio between the
concentrations of the volatile compound in the gas and liquid phase at
equilibrium [4]. Te partition coefcient is temperature dependent; the
log value of the partition coefcient is linearly related to the
temperature [5]. Temperature also has an efect on activity of enzymes
including those involved in the LOX pathway. Tomato LOX activity
increases from 0°C to 20°C, reaches its maximum activity at 20°C to
30°C and decreases at higher temperatures [6,7]. In tomatoes, HPL
activity increases from 5°C to 25°C and decreases from 25°C to 40°C
[8,9]. Tomato ADH activity increases from 10 to 20°C and decreases
from 20°C to 52°C [8,10].
Diferent fruits have diferent linoleic acid content, linolenic acid
content and LOX activity, which may explain diferent LOX volatile
profles among fruits. When extra linoleic acid was added to tomato
homogenates, the hexanal level increased [11]. Similarly, the hexenal
level was increased by addition of linolenic acid [11]. Te increase of
hexanal and 1-hexanol upon addition of linoleic acid and the increase
of (Z)-3-hexenal and (E)-2-hexenal upon addition of linolenic acid
have been reported in other studies in tomato [12,3], bell pepper [13]
and cucumber [14]. When LOX activity in tomato was decreased by
reducing expression of LOX mRNA, the levels of hexenal and hexanal
Han and Barringer, J Exp Food Chem 2016, 1:1
DOI: 10.4172/2472-0542.1000102
Research Article open access
J Exp Food Chem
ISSN:2472-0542 JEFC, an open access journal
Volume 1 • Issue 1 • 1000102
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ISSN: 2472-0542
Journal of Experimental Food
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