Short communication
A new species of Burmissus (Hemiptera: Fulgoromorpha:
Mimarachnidae) from mid-Cretaceous Burmese amber
Cihang Luo
a, b
, Tian Jiang
c
, Bo Wang
b
, Chuantao Xiao
a, *
a
School of GeoSciences, Yangtze University, Wuhan, 430100, Hubei, China
b
State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and
Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing, 210008, China
c
China University of Geosciences (Beijing), Beijing, 100083, China
article info
Article history:
Received 1 November 2019
Received in revised form
14 December 2019
Accepted in revised form 5 February 2020
Available online 11 February 2020
Keywords:
Planthoppers
Kachin amber
Mimarachnidae
New species
Burmissus szwedoi sp. nov
abstract
The eleventh species of Mimarachnidae and the second species of Burmissusis is herein described from
mid-Cretaceous Kachin amber. Burmissus szwedoi sp. nov. can be distinguished from the type species
Burmissus raunoi Shcherbakov, 2017 by the number of carinae of the mesonotum, tegmen pattern and
different broadest portion of the tegmen. Diagnostic features of the genus Burmissus are reviewed. The
phylogeny of Mimarachnidae is discussed.
© 2020 Elsevier Ltd. All rights reserved.
1. Introduction
The order Hemiptera contains the biggest number of families
among any insects, with more than three hundred extant and
extinct known families inhabiting all terrestrial and some marine
habitats (Szwedo, 2018). Fulgoromorpha (Planthoppers) is a sub-
order of the Hemiptera and subdivided into three superfamilies:
two extinct superfamilies e Permian Coleoscytoidea Martynov,
1935, Permian and Triassic Surijokocixioidea Shcherbakov, 2000
e and another superfamily Fulgoroidea Latreille, 1807 , known in
fossil records since the Jurassic (Szwedo, 2018). Up to now, various
species of Fulgoromorpha have been described from mid-
Cretaceous Burmese amber (Ross, 2019), including species from
two extant planthoppers families: Achilidae Stål, 1866 and Cixiidae
Spinola, 1839 (Cockerell, 1917; Szwedo, 2004; Brysz and Szwedo,
2018) and five extinct families: Dorytocidae Emeljanov et Shcher-
bakov, 2018 (Emeljanov and Shcherbakov, 2018), Jubisentidae
Zhang, Ren et Yao, 2019 (Zhang et al., 2019), Mimarachnidae
Shcherbakov, 2007b (Shcherbakov, 2007b), Perforissidae
Shcherbakov, 2007a (Shcherbakov, 2007a; Zhang et al., 2017), and
Yetkhatidae Song, Szwedo et Bourgoin (2019) (Song et al., 2019).
The extinct Fulgoromorpha family Mimarachnidae was placed
in the group of ‘cixiidae-like’ planthoppers (Bourgoin and Szwedo,
2008; Szwedo and Ansorge, 2015), including 9 genera and 10
species at present (Bourgoin, 2019): Mimarachne mikhailovi
Shcherbakov, 2007b and Saltissus eskovi Shcherbakov, 2007b from
Early Cretaceous deposits (ca.145-125 Ma) in Baissa, Transbaikalia,
Russia (Shcherbakov, 2007b); Nipponoridium matsuoi Fujiyama,
1978 from early Cretaceous (ca. 140-120 Ma) Kaseki-kabe locality
in Kuwajima, Japan (Szwedo, 2008); Mimamontsecia cretacea
Szwedo et Ansorge, 2015 and Chalicoridulum montsecensis Szwedo
et Ansorge, 2015 from Sierra del Montsec (Szwedo and Ansorge,
2015), north-eastern Spain of early Barremian (ca.
130.0e125.5 Ma) age; Burmissus raunoi Shcherbakov, 2017 , Dachi-
bangus trimaculatus Jiang, Szwedo et Wang, 2018, Jaculistilus oli-
gotrichus Zhang, Ren et Yao, 2018, Dachibangus formosus Fu,
Szwedo, Azar et Huang, 2019 and Mimaplax ekrypsan Jiang, Szwedo
et Wang, 2019 from mid-Cretaceous Burmese amber (Shcherbakov,
2017; Jiang et al., 2018; Zhang et al., 2018; Fu et al., 2019; Jiang et al.,
2019). Some undescribed specimens are also known from localities
like Turga (central Siberia) of early Cretaceous, Khurilt (Mongolia)
of Barremian or Aptian, Khetana (East Siberia) of middle Albian, and * Corresponding author.
E-mail address: ctxiao@yangtzeu.edu.cn (C. Xiao).
Contents lists available at ScienceDirect
Cretaceous Research
journal homepage: www.elsevier.com/locate/CretRes
https://doi.org/10.1016/j.cretres.2020.104417
0195-6671/© 2020 Elsevier Ltd. All rights reserved.
Cretaceous Research 110 (2020) 104417