ORIGINAL PAPER
Med Mol Morphol (2009) 42:212–215 © The Japanese Society for Clinical Molecular Morphology 2009
DOI 10.1007/s00795-009-0464-9
Masato Saitoh · Yoshito Kurashige · Michiko Nishimura
Mami Yamazaki · Seiji Igarashi · Tohru Kaku
Yoshihiro Abiko
Expression of claudin-4 and -7 in porcine gingival junctional epithelium
Abstract Junctional epithelium, a nonkeratinized stratified
epithelium, extends apically in apposition to the surface of
the enamel to form a seal between the epithelium and the
tooth. Desmosomes and gap junctions adhere to the junc-
tional epithelium through cell–cell contact, but no evidence
of tight junctions has been found. Recently, tight junction
hallmark proteins and tight junction-related structures have
been identified in stratified squamous epithelium. The
present study examined whether tight junction proteins
were expressed in the junctional epithelium. We used
immunohistochemical techniques to observe expression of
claudin-1, -4, -5, -7, and occludin in porcine gingival junc-
tional epithelium. Claudin-4 exhibited immunoreactivity in
the intercellular spaces of all layers of the oral epithelium
and the junctional epithelium. Stronger expression was
observed in junctional epithelial cells adjacent to the inner
and outer basal laminae than in the inner cell layers. Immu-
nohistochemical positivity for claudin-7 was clearly observed
in the junctional epithelium, but only a faint positivity was
observed in the basal layer of the oral epithelium. No immu-
nohistochemical positivity for claudin-1, -5, or occludin was
observed in the junctional epithelium. RT-PCR assay con-
firmed expression of porcine claudin-4 and -7 mRNAs in
the junctional epithelium. These findings indicate that
M. Saitoh
Department of Dental Science, Institute of Personalized Medical
Science, Hokkaido, Japan
M. Saitoh · Y. Kurashige · S. Igarashi
Division of Pediatric Dentistry, Department of Oral Growth and
Development, Health Sciences University of Hokkaido, Hokkaido,
Japan
M. Nishimura · M. Yamazaki · T. Kaku · Y. Abiko
Division of Clinical Oral Pathology, Department of Human Biology
and Pathophysiology, School of Dentistry, Health Sciences
University of Hokkaido, Hokkaido, Japan
Y. Abiko (*)
Division of Oral Medicine and Pathology, Department of Dental
Science, Institute of Personalized Medical Science, Health Sciences
University of Hokkaido, 2-5 Ainosato, Kita-ku, Sapporo, Hokkaido
002-8072, Japan
Tel. +81-11-778-7557; Fax +81-11-770-5034
e-mail: yoshi-ab@hoku-iryo-u.ac.jp
claudin-4 and -7 may play a role in the junctional epithelium
even in the absence of tight junctions.
Key words Claudin · Tight junction · Stratified
squamous epithelium · Oral gingival epithelium · Junctional
epithelium
Introduction
The gingiva is located around the neck of each tooth and
resists the forces of mastication. It is composed of gingival
epithelium and gingival connective tissue. The gingival epi-
thelium is classified into gingival oral epithelium, sulcular
epithelium, and junctional epithelium. The junctional epi-
thelium, a nonkeratinized epithelium, forms the floor of the
gingival sulcus and extends apically in apposition to the
surface of the enamel to form a seal between the epithelium
and the tooth. The junctional epithelium is characterized by
wide intercellular spaces as a result of the reduced number
of desmosomes. The intercellular spaces contain varying
numbers of neutrophils. In addition to the desmosomes, gap
junctions are formed by cell–cell contact of the junctional
epithelium.
1
A further type of intercellular junction, the
tight junction, is considered less important in stratified
squamous epithelia such as epidermis, oral epithelium, and
junctional epithelium because the abundance of desmo-
somes mediates a firm mechanical stability between the
cells. Tight junctions are composed of three major trans-
membrane proteins, namely claudin, occludin, and junc-
tional adhesion molecules.
2–4
Claudins fulfill a barrier
function in epithelial and endothelial tissues as they are
thought to be responsible for the variety of electrical resis-
tance and paracellular ionic selectivity seen in epithelia and
endothelia.
5–7
The claudin family consists of 24 members,
and four transmembrane domains have been identified in
different species. The claudins show specific patterns of
expression in different tissues.
8
Occludin may be more
important for cell signaling than for forming the paracellu-
lar barrier.
9,10
Recent reports have demonstrated localiza-
Received: April 30, 2009 / Accepted: July 17, 2009