Citation: Hotopp, I.; Walther, B.;
Fuelling, O.; Reil, D.; Hesse, C.;
Below, D.A.; Imholt, C.; Jacob, J.
Habitat and Season Effects on Small
Mammal Bycatch in Live Trapping.
Biology 2022, 11, 1806. https://
doi.org/10.3390/biology11121806
Academic Editor: Carles Vilà
Received: 18 November 2022
Accepted: 7 December 2022
Published: 13 December 2022
Publisher’s Note: MDPI stays neutral
with regard to jurisdictional claims in
published maps and institutional affil-
iations.
Copyright: © 2022 by the authors.
Licensee MDPI, Basel, Switzerland.
This article is an open access article
distributed under the terms and
conditions of the Creative Commons
Attribution (CC BY) license (https://
creativecommons.org/licenses/by/
4.0/).
biology
Article
Habitat and Season Effects on Small Mammal Bycatch in
Live Trapping
Ines Hotopp
1
, Bernd Walther
2
, Olaf Fuelling
1
, Daniela Reil
2
, Christin Hesse
2
, Diana Alexandra Below
2
,
Christian Imholt
2
and Jens Jacob
2,
*
1
tier3 Solutions GmbH, Kolberger Str. 61–63, 51381 Leverkusen, Germany
2
Rodent Research, Institute for Epidemiology and Pathogen Diagnostics, Julius Kühn-Institute (JKI)–Federal
Research Centre for Cultivated Plants, Toppheideweg 88, 48161 Münster, Germany
* Correspondence: jens.jacob@julius-kuehn.de
Simple Summary: Trapping particular small mammal species is frequently used for scientific pur-
poses but unnecessary bycatch can occur. Live trapping conducted over the last decade in Ger-
many using Ugglan multiple capture traps in grassland, forest and margin habitats revealed about
30% bycatch when target species were common voles (Microtus arvalis) in grassland and common
voles and bank voles (Clethrionomys glareolus) in margins and forests. This was more pronounced
in spring and along margins. Species mentioned on the early warning list according to the Red List
Germany were higher in numbers and proportion in spring and in grassland. The results of the study
will help to avoid periods with enhanced presence of bycatch including endangered species (if the
purpose of the study allows) or to pay particular attention in certain seasons and habitats when the
occurrence of bycatch is most likely.
Abstract: Trapping small mammals is frequently used to study the dynamics, demography, behavior
and presence of pathogens. When only particular small mammal species are in the focus of interest,
all other species are unnecessary bycatch. We analyzed data from extensive live trapping campaigns
conducted over the last decade in Germany, following a consistent standard trapping protocol that
resulted in about 18,500 captures of small mammals. Animals were trapped with Ugglan multiple
capture traps in grassland, forest and margin habitat. Trap success and the proportion of bycatch
were about 30% when target species were common voles (Microtus arvalis) in grassland and common
voles and bank voles (Clethrionomys glareolus) in margins and forests. This was more pronounced
in spring and along margins. Species mentioned in the early warning list according to the Red List
Germany were higher in numbers and proportion in spring and in grassland. The results will help to
avoid periods with enhanced presence of bycatch, including endangered species (if the purpose of
the study allows) or to pay particular attention in certain seasons and habitats when the occurrence
of bycatch is most likely.
Keywords: conservation; non-target species; endangered species; rodents; Ugglan traps; voles
1. Introduction
Capturing wildlife is important to humans as wild animals are recognised as a source
of nutrients and material for shelter, tools, medicine and cultural objects, etc. For millennia,
wild animals have been vital in the process of domestication, for support in hunting and
herding, lifting and transporting objects and other work, religious rites and as guards,
as well as companions. The importance of wildlife in these aspects was later supple-
mented by the relevance of wildlife in research. In many cases, the species required needs
to be obtained.
While capture for food does not necessarily require the animal to stay alive, their use
for observational studies does. In some aspects of research such as the study of animal
Biology 2022, 11, 1806. https://doi.org/10.3390/biology11121806 https://www.mdpi.com/journal/biology