Citation: Tsotetsi, T.; Nephali, L.;
Malebe, M.; Tugizimana, F. Bacillus
for Plant Growth Promotion and
Stress Resilience: What Have We
Learned? Plants 2022, 11, 2482.
https://doi.org/10.3390/
plants11192482
Academic Editor: Barbara
Hawrylak-Nowak
Received: 3 September 2022
Accepted: 19 September 2022
Published: 22 September 2022
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plants
Review
Bacillus for Plant Growth Promotion and Stress Resilience:
What Have We Learned?
Teboho Tsotetsi
1,
*, Lerato Nephali
1
, Motumiseng Malebe
1
and Fidele Tugizimana
1,2,
*
1
Department of Biochemistry, University of Johannesburg, Auckland Park, Johannesburg 2006, South Africa
2
International R&D Division, Omnia Nutriology, Omnia Group (Pty) Ltd., Johannesburg 2021, South Africa
* Correspondence: ttsotetsi@uj.ac.za (T.T.); fidele.tugizimana@omnia.co.za (F.T.); Tel.: +27-11-559-7784 (F.T.);
Fax: +27-11-559-2370 (F.T.)
Abstract: The rhizosphere is a thin film of soil that surrounds plant roots and the primary location
of nutrient uptake, and is where important physiological, chemical, and biological activities are
occurring. Many microbes invade the rhizosphere and have the capacity to promote plant growth
and health. Bacillus spp. is the most prominent plant growth promoting rhizobacteria due to its
ability to form long-lived, stress-tolerant spores. Bacillus-plant interactions are driven by chemical
languages constructed by a wide spectrum of metabolites and lead to enhanced plant growth and
defenses. Thus, this review is a synthesis and a critical assessment of the current literature on the
application of Bacillus spp. in agriculture, highlighting gaps that remain to be explored to improve
and expand on the Bacillus-based biostimulants. Furthermore, we suggest that omics sciences, with a
focus on metabolomics, offer unique opportunities to illuminate the chemical intercommunications
between Bacillus and plants, to elucidate biochemical and molecular details on modes of action of
Bacillus-based formulations, to generate more actionable insights on cellular and molecular events
that explain the Bacillus-induced growth promotion and stress resilience in plants.
Keywords: Bacillus; biostimulant; plant growth; stress resilience; metabolomics
1. Introduction
Edaphic factors and genetics play a pivotal role in the growth and yield of crop
plants [1]. Over the past decades, genetic engineering and plant breeding approaches
have been employed to develop new cultivars with desired traits, such as high yield
and resistance to environmental stresses [2]. However, there is a less commercial success
for genetically modified crops due to ethical constraints concerning genetically modified
organisms (GMO) [3]. To obtain better crop yield, applications of chemical fertilizers have
been the opted strategy. However, over time, studies and empirical evidence have shown
that this traditional method—the use of chemical fertilizers—is not sustainable due to the
inherent negative effects these products have on the environment. The excessive utilization
of chemical fertilizers has shown to lead to toxic build-up of heavy metals, soil acidification
and soil crust, thereby reducing the soil content of organic matter and humic substance.
Soil acidification reduces crop phosphate intake, raises the concentration of harmful ions in
the soil, and inhibits crop growth [4].
The incorporation of biostimulants, such as plant growth-promoting rhizobacteria
(PGPR)-based formulations, in cropping systems has increasingly shown to be a promising
strategy for sustainable agriculture and global food security, aligning with the United
Nations sustainable development goals (SDGs) [5]. A broad array of bacterial species has
been reported to possess plant growth-promoting attributes with the prominent species be-
longing to the genus Bacillus. Members of the genus Bacillus are ubiquitous, Gram-positive,
and aerobic bacteria [6,7]. Bacillus species produce a multitude of enzymes, antibiotics, and
metabolites which give them prominent applications in various sectors such as pharmaceu-
ticals and agriculture. Furthermore, their uniqueness and popularity arise from their spore
Plants 2022, 11, 2482. https://doi.org/10.3390/plants11192482 https://www.mdpi.com/journal/plants