Romanian Biotechnological Letters Vol. 17, No.3, 2012
Copyright © 2012 University of Bucharest Printed in Romania. All rights reserved
ORIGINAL PAPER
7227
Effects of long-term salt stress on antioxidant system, chlorophyll and
proline contents in pea leaves
Received for publication, January 10, 2010
Accepted, June 11, 2011
LOKMAN OZTURK
1
, YAVUZ DEMIR*
2
, ALI UNLUKARA
3
, ILHAMI KARATAS
1
,
AHMET KURUNC
4
AND ORAL DUZDEMIR
5
1
Department of Biology, Faculty of Arts and Sciences, Gaziosmanpasa University, 60250-
Tokat, Turkey
2
Department of Biology, Faculty of Education, Atatürk University, 25240-Erzurum, Turkey.
3
Department of Agricultural Structures and Irrigation, Faculty of Agriculture, Erciyes
University, 38039-Kayseri, Turkey
4
Department of Agricultural Structures and Irrigation, Faculty of Agriculture, Akdeniz
University, 07058-Antalya, Turkey
5
Vocational School, Karatekin University, 18100-Çankırı, Turkey
Abstract
The effects of long-term salt stress on the contents of chlorophyll, proline, protein, hydrogen
peroxide (H
2
O
2
), and malondialdehyde (MDA) in terms of lipid peroxidation, and on the changes in
activities of antioxidant enzymes such as superoxide dismutase (SOD), catalase (CAT), ascorbate
peroxidase (APX) and peroxidase (POD) in leaves of pea (Pisum sativum cv Rona) were investigated
in field conditions. Salinity and irrigation water regime experiments were set up in pots as
randomized plot experimental designs with five replications. The experiment focused on pea
responses to irrigation water salinity by irrigating the crops using 6 different levels of saline water
(0.7, 2.0, 3.0, 4.0, 5.0 and 7.0 dS m
-1
) with a constant leaching fraction (LF = 0.30). The pots were
located in the experimental area under a polyethylene cover which was about 1.8 m height from the
surface for the purpose of eliminating rainfall effect on the experiments. Chlorophyll and protein
contents were significantly decreased while accumulation of proline was enhanced with increased
electrical conductivity (EC). Both MDA and H
2
O
2
contents were reduced in the result of high salts
application. All of the salt treatments increased total SOD activity significantly, as a remarkable
increase in POD activity was observed especially at 5.0 and 7.0 dS/m EC. CAT and APX activities
generally decreased in salt stressed seedlings. Our study indicates that its acquisition of salt
tolerance may be a consequence of improved resistance to oxidative stress via increased activities of
peroxidase and the superoxide dismutase/ascorbate-glutathione cycle.
Key words: Antioxidant Enzyme, Pea, Proline, Salinity, MDA, H
2
O
2
.
Abbrevations: EDTA, Ethylenediaminetetraacetic acid; PVPP, polyvinyl polypirrolidone,
Introduction
Salinity in the soil and irrigation water is an environmental problem and a major
constraint for crop production. Salt stress (NaCI) in plants influences some basic plant
metabolic processes such as, photosynthesis, protein synthesis, and energy and lipid
metabolism [1]. Salt stress (NaCl) has both osmotic (cell dehydration) and toxic (ion
accumulation) effects on plant cells, impairing growth, ion homeostasis, photosynthesis and
nitrogen fixation among other key physiological processes. Nowadays, several markers of
stress have been measured to assess the physiological status of the plants [1]. These include
proline and soluble protein contents, photosynthetic pigments and metabolic products of
oxidative damage. In addition to other environmental stresses, salt stress also causes oxidative
damage, thereby affecting cellular membrane integrity, enzyme activities, and functioning of