Acta Tropica 240 (2023) 106858
Available online 5 February 2023
0001-706X/© 2023 Elsevier B.V. All rights reserved.
Neem-based products as potential eco-friendly mosquito control agents
over conventional eco-toxic chemical pesticides-A review
Soumendranath Chatterjee
a, *
, Souvik Bag
a
, Debraj Biswal
b
, Dipanwita Sarkar Paria
c
,
Raktima Bandyopadhyay
d
, Basanta Sarkar
a
, Abhijit Mandal
a
, Tushar Kanti Dangar
e
a
Parasitology and Microbiology Research Laboratory, Department of Zoology, The University of Burdwan, Burdwan 713104, West Bengal, India
b
Department of Zoology, Government General Degree College at Mangalkote, Burdwan 713132, West Bengal, India
c
Department of Zoology, Chandernagore College, Hooghly 712136, West Bengal, India
d
Department of Nutrition, AKPC Mahavidyalaya, Hooghly 712611, West Bengal, India
e
Microbiology Laboratory, Division of Crop Production, National Rice Research Institute, Cuttack 753006, Odisha, India
A R T I C L E INFO
Keywords:
Arbovirus vector
Azadirachta indica
Larvicide
Mosquito repellent
Neem oil formulations
Eco-friendly mosquito control
ABSTRACT
Mosquitoes cause serious health hazards for millions of people across the globe by acting as vectors of deadly
communicable diseases like malaria, flariasis, dengue and yellow fever. Use of conventional chemical in-
secticides to control mosquito vectors has led to the development of biological resistance in them along with
adverse environmental consequences. In this light, the recent years have witnessed enormous efforts of re-
searchers to develop eco-friendly and cost-effective alternatives with special emphasis on plant-derived mos-
quitocidal compounds. Neem oil, derived from neem seeds (Azadirachta indica A. Juss, Meliaceae), has been
proved to be an excellent candidate against a wide range of vectors of medical and veterinary importance
including mosquitoes. It is environment-friendly, and target-specifc at the same time. The active ingredients of
neem oil include limonoids like azadirachtin A, nimbin, salannin and numerous other substances that are still
waiting to be discovered. Of these, azadirachtin has been shown to be very effective and is mainly responsible for
its toxic effects. The quality of the neem oil depends on its azadirachtin content which, in turn, depends on its
manufacturing process. Neem oil can be used directly or as nanoemulsions or nanoparticles or even in the form of
effervescent tablets. When added to natural breeding habitat waters they exert their mosquitocidal effects by
acting as ovicides, larvicides, pupicides and/or oviposition repellents. The effects are generated by impairing the
physiological pathways of the immature stages of mosquitoes or directly by causing physical deformities that
impede their development. Neem oil when used directly has certain disadvantages mainly related to its disin-
tegration under atmospheric conditions rendering it ineffective. However, many of its formulations have been
reported to remain stable under environmental conditions retaining its effciency for a long time. Similarly, neem
seed cake has also been found to be effective against the mosquito vectors. The greatest advantage is that the
target species do not develop resistance against neem-based products mainly because of the innumerable number
of chemicals present in neem and their combinations. This makes neem-based products highly potential yet
unexplored candidates of mosquito control agents. The current review helps to elucidate the roles of neem oil and
its various derivatives on mosquito vectors of public health concern.
1. Introduction
Mosquitoes (Diptera: Culicidae) transmit deadly human diseases like
malaria, dengue haemorrhagic fever, flariasis, Japanese encephalitis
and yellow fever, making them one of the leading causes of human
mortality (Mittal, 2003). According to records, mosquito-borne illnesses
cause death of about one million people annually (World Health
Organization, 2012). Under this situation, vector control seems to be the
most effective strategy to reduce the number of deaths. The mainstays of
mosquito vector control include the use of larvicidal chemicals and
application of repellents or insecticide-treated bed nets to keep away the
adults. For instance, organophosphates and insect growth regulators are
primarily used to target the Culicid (Family: Culicidae) larvae. However,
widespread use of chemical pesticides has been associated with the
* Corresponding author.
E-mail address: soumen.microbiology@gmail.com (S. Chatterjee).
Contents lists available at ScienceDirect
Acta Tropica
journal homepage: www.elsevier.com/locate/actatropica
https://doi.org/10.1016/j.actatropica.2023.106858
Received 1 December 2022; Received in revised form 2 February 2023; Accepted 3 February 2023