Vegetatio 75: 73-80, 1988
© Kluwer Academic Publishers, Dordrecht - Printed in the Netherlands 73
Long-term vegetation changes on the stabilized dunes of Dofiana National
Park (SW Spain)
Manuel Granados Corona, Angel Martin Vicente & Francisco Garcia Novo
Department of Ecology, University of Sevilla, Ap. 1095, 41080 Sevilla, Spain
Accepted 10.3.1988
Keywords: Coastal dune, Human impact, Mediterranean forest, Mediterranean scrub
Abstract
Changes in the vegetation of stabilized sand dunes of Dofiana National Park in the last four centuries have
been analysed, combining ecological and historical sources of information.
Quantitative vegetation data have been obtained from 5 inventories of a 15 km long transect, consisting of
34 points where nearby woody plants have been recorded. Inventories were taken in 1636, 1647, 1652, 1682
and 1859. The analysis of these data, combined with climatic and management information on the area, reveals
that until 1636 a mediterranean forest dominated by Quercus suber, existed; it was used as a game preserve.
From 1636 to 1682 vegetation underwent profound changes with a marked drop in trees and mature scrub.
The vegetation was replaced by a pioneer scrub species. This rapid change was brought about by the introduc-
tion of cattle and pigs, timber and cork exploitation, and the use of prescribed fire. The climatic crisis that
arose circa 1700 led to erosion of the sandy soils, which made the vegetation change irreversibly. Livestock
pressure hampered natural forest regeneration until 1737 when Pinuspinea plantations started. From then on-
wards little change in vegetation is known to have occurred.
It is suggested that the main steps of change in Dofiana vegetation history may have occurred in many other
areas of the Mediterranean Basin.
Nomenclature: follows Tutin et aL (1964-1980), Flora Europaea.
Introduction
The Dofiana National Park (Reserve of the Bi-
osphere of the MaB Program; 50720 ha) is situated
on the SW coast of Spain between 3707 ' and
36°48'N and 6°12 ' and 6°34'W. The climate is of
a mediterranean subhumid type with some atlantic
influence. According to Gaussen (1949) it can be
classified as thermo-mediterranean. Average rain-
fall is 620 mm with less than 75 mm from May to
September. The average daily mean temperature is
9.3 °C during the coldest months (January and De-
cember). The summer temperature is high with a
daily mean of 23.9 °C for July, the warmest month.
Due to their origin as fixed holocene sand dunes, the
stabilized sands have an undulating topography
(Vanney & Menanteau 1985). The presence of a
hanging water table along with the permeability of
the sandy substrate, makes higher zones markedly
arid in summer, while depressions are humid with
occasional flooding in winter.
The present vegetation is dominated by mediterra-
nean scrub, the floristic composition of which close-
ly follows local topography. On higher places an