Original article
a-Aminoalkylphosphonates as a tool in experimental optimisation
of P1 side chain shape of potential inhibitors in S1 pocket of leucine-
and neutral aminopeptidases
Marcin DrTg
a,
*, Jolanta Grembecka
a
, Malgorzata Pawelczak
b
, Pawel Kafarski
a,
*
a
Institute of Organic Chemistry, Biochemistry and Biotechnology, University of Technology, Wybrzeze Wyspianskiego 27, 50-370 Wroclaw, Poland
b
Faculty of Chemistry, University of Opole, Opole, Oleska 48, Poland
Received 26 October 2004; received in revised form 14 February 2005; accepted 24 February 2005
Available online 20 April 2005
Abstract
The synthesis and biological activity studies of the series of structurally different a-aminoalkylphosphonates were performed in order to
optimise the shape of the side chain of the potential inhibitors in S1 pocket of leucine aminopeptidase [E.C.3.4.11.1]. Analysis of a series of
compounds with aromatic, aliphatic and alicyclic P1 side chains enabled to find out the structural features, optimal for that fragment of
inhibitors of LAP. The most active among all investigated compounds were the phosphonic analogues of homo-tyrosine (K
i
= 120 nM) and
homo-phenylalanine (K
i
= 140 nM), which even as racemic mixtures were better inhibitors in comparison with the best till now-phosphonic
analogue of L-leucine (230 nM). Additional comparison of the inhibitory activity obtained for aminopeptidase N (APN, E.C.3.4.11.2) give
insight into structural preferences of both enzymes.
© 2005 Elsevier SAS. All rights reserved.
Keywords: Aminophosphonates; Inhibitor; Leucine aminopeptidase; Aminopeptidase N
1. Introduction
Aminopeptidases are enzymes that catalyse the removal
of amino acids from the N-terminus of a peptide or protein.
These enzymes have been found in bacteria, yeast, plants,
animal and human tissues, what is considerably ascribed to
their broad substrate specificity [1]. They play such a biologi-
cal functions as protein maturation, activation and degrada-
tion of bioactive peptides, hormone level regulation and con-
trol of cell proliferation [2,3]. It is well known, that elevated
concentration of these enzymes was observed in such patho-
logical disorders as cancer and cataracts [1,3]. In the case of
the carcinogenesis, the methionine aminopeptidases has been
shown to be a promising target for development of anti-
angiogenesis agents [4]. Additionally, the importance of these
enzymes arises from the observation, that some of them might
play the role in the apoptosis of cancer cells, what makes
them interesting targets in oncological research (with methion-
ine and neutral aminopeptidases being the most promising
targets) [5].
Aminopeptidases can be subdivided into two major groups,
with the first one being responsible for the hydrolysis of amino
acids with hydrophobic side chains from N-terminus of the
peptide or protein, and the second, which specifically remove
other amino acid residues [3].
Within the first group, leucine aminopeptidase (LAP,
E.C.3.4.11.1) is one of the most detailed characterised enzyme
with respect to its sequence, structure and mechanism of
action. Human LAP plays an important role in the early stages
of HIV infection, where the elevated concentration of this
enzyme was observed [6]. The unusual activity of this enzyme
has also been observed in such pathological disorders as can-
cer, inflammation of liver or eye cataract [7–9]. Because of
this leucine aminopeptidase is often called the stress enzyme.
Substantial insight into the structure of the LAP derives
from X-ray structures of the enzyme from bovine lens deter-
mined in its native form and complexed with transition state
inhibitors such as: bestatin, amastatin, phosphonic analogue
of L-leucine and L-leucinal coordinated to the active site.
* Corresponding author. Tel.: +48 71 320 4027; fax: +48 71 328 4064.
E-mail address: marcin.drag@pwr.wroc.pl (M. DrTg).
European Journal of Medicinal Chemistry 40 (2005) 764–771
www.elsevier.com/locate/ejmech
0223-5234/$ - see front matter © 2005 Elsevier SAS. All rights reserved.
doi:10.1016/j.ejmech.2005.02.011