              Mohamed Ali Bouaziz 1* , Fatma Abbes 1 , Abir Mokni 1 , Christophe Blecker 2 , Hamadi Attia 1 and Souhail Besbes 1               ! "#"$  % & ’  ( ) * *+ ’  %,   -  .- & /#"# + M.A BOUAZIZ et al. *      E%MAIL: medalibouaziz@yahoo.fr TEL: +216.74.274.088; FAX: +216.74.675.761; This article has been accepted for publication and undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process which may lead to differences between this version and the Version of Record. Please cite this article as an ‘Accepted Article’, doi: 10.1111/jtxs.12225 This article is protected by copyright. All rights reserved.