Citation: Ashi, H.; Almalki, M.H.K.;
Hamed, E.A.; Ramadan, W.S.;
Alahmadi, T.F.H.; Alami, O.T.; Arafa,
S.H.; Alshareef, A.K.; Alsulami, F.S.;
Alharbi, A.F.; et al. Protective and
Therapeutic Effects of Lactic Acid
Bacteria against Aflatoxin B1 Toxicity
to Rat Organs. Microorganisms 2023,
11, 1703. https://doi.org/10.3390/
microorganisms11071703
Academic Editor: Salam A. Ibrahim
Received: 11 May 2023
Revised: 25 June 2023
Accepted: 26 June 2023
Published: 29 June 2023
Copyright: © 2023 by the authors.
Licensee MDPI, Basel, Switzerland.
This article is an open access article
distributed under the terms and
conditions of the Creative Commons
Attribution (CC BY) license (https://
creativecommons.org/licenses/by/
4.0/).
microorganisms
Article
Protective and Therapeutic Effects of Lactic Acid Bacteria
against Aflatoxin B1 Toxicity to Rat Organs
Hayat Ashi
1,2
, Meshal H. K. Almalki
1,2
, Enas A. Hamed
3
, Wafaa S. Ramadan
4
, Tahani F. H. Alahmadi
1,2
,
Outour Tariq Alami
1,2
, Sara H. Arafa
1,2
, Atheer K. Alshareef
1,2
, Fatimah S. Alsulami
1,2
, Areej F. Alharbi
1,2
,
Manahil S. Al-Harbi
1,2
, Ebtehal H. Alqurashi
1,2
, Shirin Aashi
5
, Youssef A. Alzahrani
5
, Khaled Elbanna
1,2,6
and Hussein H. Abulreesh
1,2,
*
1
Department of Biology, Faculty of Applied Science, Umm Al-Qura University, Makkah 21955, Saudi Arabia;
dr.ht.111@hotmail.com (H.A.); mhmaleki@uqu.edu.sa (M.H.K.A.); tootaa.2030@gmail.com (T.F.H.A.);
outour-alami@hotmail.com (O.T.A.); arafa.sa@hotmail.com (S.H.A.); alal-2021@hotmail.com (A.K.A.);
fatima.alsulami@hotmail.com (F.S.A.); areej.f.alharbi@hotmail.com (A.F.A.);
s44285749@st.uqu.edu.sa (M.S.A.-H.); ebtehal373@yahoo.com (E.H.A.); kab00@fayoum.edu.eg (K.E.)
2
Research Laboratories Unit, Faculty of Applied Science, Umm Al-Qura University, Makkah 21955, Saudi Arabia
3
Department of Medical Physiology, Faculty of Medicine, Assiut University, Assiut 71515, Egypt;
eah3a2010@aun.edu.eg
4
Department of Anatomy, Faculty of Medicine, King Abdulaziz University, Jeddah 21589, Saudi Arabia;
wramadhan@kau.edu.sa
5
College of Pharmacy, King Abdulaziz University, Jeddah 21589, Saudi Arabia;
sheren_141@hotmail.com (S.A.); youssefalzahrani18@gmail.com (Y.A.A.)
6
Department of Agricultural Microbiology, Faculty of Agriculture, Fayoum University, Fayoum 63514, Egypt
* Correspondence: hhabulreesh@uqu.edu.sa
Abstract: Background: Aflatoxin (AF), a metabolite of Aspergillus flavus, is injurious to vital body
organs. The bacterial defense against such mycotoxins has attracted significant attention. Lactic acid
bacteria (LAB) are known to ameliorate AF toxicity. Methods: Thirty adult male rats were divided
into six groups (five each) to perform the experiments. The control (Co) group was fed a basal diet
and water. Each of the following periods lasted 21 days: the milk (MK) group orally received milk
(500 μL); LAB suspension (500 μL) containing 10
7
cfu/mL was orally provided to the LAB group; AF
(0.5 mg/kg) was orally given to the AF group; and a combination of AF and LAB was administered
to the AF + LAB group. The AF/LAB group was initially given AF for 21 days, followed by LAB
for the same period. Finally, the rats were dissected to retrieve blood and tissue samples for hemato-
logical, biochemical, and histological studies. Results: The results revealed a significant decrease in
RBCs, lymphocytes, total proteins, eosinophil count, albumin, and uric acid, whereas the levels of
WBCs, monocytes, neutrophils, creatinine, urea, aspartate aminotransferase, alkaline phosphatase,
alanine aminotransferase, lactate dehydrogenase, and creatinine kinase significantly increased in
the AF group in comparison to the control group. The histological examination of the AF group
revealed necrosis and apoptosis of the kidney’s glomeruli and renal tubules, nuclei vacuolization
and apoptosis of hepatocytes, congestion of the liver’s dilated portal vein, lymphoid depletion in the
white pulp, localized hemorrhages, hemosiderin pigment deposition in the spleen, and vacuolization
of seminiferous tubules with a complete loss of testis spermatogenic cells. Meanwhile, protective and
therapeutic LAB administration in AF-treated rats improved the hematological, biochemical, and
histological changes. Conclusions: The study revealed LAB-based amelioration to AFB1-induced
disruptions of the kidney, liver, spleen, and testis by inhibiting tissue damage. The therapeutic effects
of LAB were comparatively more pronounced than the protective effects.
Keywords: aflatoxin; lactic acid bacteria; liver; kidney; protective; rats; spleen; structure; testes; therapeutic
Microorganisms 2023, 11, 1703. https://doi.org/10.3390/microorganisms11071703 https://www.mdpi.com/journal/microorganisms