International Journal of Poultry Science 10 (4): 257-265, 2011
ISSN 1682-8356
© Asian Network for Scientific Information, 2011
Corresponding Author: Samuel Besong, Department of Human Ecology, Delaware State University, 1200 North DuPont HWY, John
Price Bldg RM #103, Dover, DE 19901
257
Effect of Njangsa Oil Seeds (Ricinodendron heudelitii) on Expression of
Stearoyl-coenzyme a Desaturase in Laying Hens
Samuel Besong , Celestine Fosung , Ayasha Jones and Vincent Fondong
1 1 1,2 2
Department of Human Ecology, Delaware State University, 1200 North DuPont HWY,
1
John Price Bldg RM #103, Dover, DE 19901
Department of Biological Science, Delaware State University,
2
1200 North DuPont HWY, Dover, DE 19901
Abstract: Pathogenesis of liver steatosis in laying hen remains poorly understood. However, it has been
suggested that regulation of the expression of Stearoyl-Coenzyme A Desaturase-1 (SCD-1) may alter hepatic
lipid metabolism and prevent liver steatosis in laying hens. SCD-1 is an integral membrane protein that
catalyzes the biosynthesis of monounsaturated fatty acids required for the synthesis of triglycerides,
cholesterol esters and phospholipids. This study was conducted to determine whether Njangsa Oil Seed
(NOS) (Ricinodendron heudelotii) which is rich in polyunsaturated fatty acids, specifically omega-3 fatty acids
can regulate the expression of SCD-1 gene and lower hepatic lipid accumulation in laying hens. The
experiment was carried out with seventy-two white leghorn hens (24 months old) which were randomly
assigned to three layer diet supplemented with 0%, 10% or 20% NOS for 6 weeks. The mRNA levels in SCD-
1 gene was quantified using forward and reverse primers specific to chicken SCD-1 in real-time reverse
transcription-PCR (RT-PCR) technique with SYBER Green detection. Histological studies with liver tissues
were performed with oil red stain and electron transmission microscope. Results indicate that NOS
supplementation at 10% level tended to increase (p>0.05) SCD-1 mRNA expression in hens. Histological
examination revealed micro-vesicular fat accumulation in the liver of hen fed NOS diets, indicating that
supplementation of NOS in the diet may not prevent development of fatty liver. Results from this study
suggest that NOS supplementation may regulate genes involve in lipid biosynthesis but may not prevent liver
steatosis.
Key words: Njangsa oil seeds, stearoyl-coenzyme A desaturase-1, mRNA, laying hen, steatosis
INTRODUCTION
Fatty Liver Syndrome (FLS) is one of the most common
metabolic disorders seen in laying hens in the period of
high egg production. Liver cells distended with fat
vacuoles and different size of hemorrhages (Riddell,
1997). Some studies have reported that the fat content of
liver in the hen diagnosed with hepatic steatosis ranges
from 40-70% dry weight (Riddell, 1997). Excess fat
impairs liver functions and alters carbohydrate and
protein metabolism and markedly impair glycogenesis
and gluconeogenesis (Felig et al., 1970). Although the
real cause of fatty liver disease in laying hens is not well
understood, however excessive accumulation of lipids
predominantly triacylglycerols in hepatocytes occur when
VLDL transport is disrupted.
Triacylglycerol secretion in the form of VLDL from liver
cells is highly dependent on the activity of stearoyl- CoA
desaturase, which converts saturated fatty acids into
monounsaturated fatty acids, specifically oleic acids
(Cook, 1991). Hepatic packaging and secretion of VLDL
require synthesis of apoB-100 as well as sufficient
amount of oleic acid (C18:1, n-9), originating from a diet
or synthesized by stearoyl-CoA desaturase (Ntambi,
1999). Recent studies with mouse model have reported
that SCD1-deficient (SCD1 ) mice showed reduced
–/–
synthesis of lipids, especially triglycerides (Miyazaki et
al., 2001). SCD1 mice have low levels of triglycerides
–/–
in VLDL and increased plasma HDL-cholesterol levels
(Cohen et al., 2002; Huuskonen et al., 2001). These
observations suggest that regulating SCD-1 gene would
be a novel intervention strategy to improve poultry health.
Thus, we hypothesized that incorporating njangsa oil
seed (Ricinodendron heudelotii), rich in omega-3 fatty
acids in poultry ration would reduce hepatic triglyceride
accumulation through its action on the expression of
SCD-1 genes. Njangsa (Ricinodendron heudelotii;
family Euphorbiaceae), a tropical shrub from which oil
seeds are harvested, grows or cultivated in West and
Central Africa-specifically in Cameroon. Ricinodendron
heudelotii reaches maturity between 4-5 years before
producing fruits. The fruits are usually manually shelled
to collect the seeds and dried. The seeds are grinded
and used for soup and as an ingredient for seasoning
baked meats and fish. The oil seed is an economical