Detection of amoeba and amoeba-associated microorganisms (AAMs) from
natural and hospital environments
M. F. Caeiro*, R. Costa *
,
**
,
*****, A. Amorim**, F. F. Vale***, S. Ferreira****, F. Morgado****
and A. P. Alves de Matos*
,
*****
* Centro de Estudos do Ambiente e do Mar (CESAM), Faculdade de Ciências da Universidade de
Lisboa (FCUL), Campo Grande, 1749-016, Lisboa
** Centro de Oceanografia, FCUL, Campo Grande, 1149-016, Lisboa
*** Faculdade de Engenharia, Universidade Católica Portuguesa, 2635-631 Rio de Mouro
**** CESAM, Universidade de Aveiro, Campus Universitário de Santiago, 3810-193 Aveiro
***** Centro Hospitalar de Lisboa Central - HCC, Anatomia Patológica, R. da Beneficência 8, 1069-
166 Lisboa
Amoebae graze on bacteria from the environment. In some cases, bacteria acquire resistance to the
destructive action of amoebae and survive inside the host [1]. The resistance of microorganisms to
amoebae is often accompanied by resistance to the physiologically related human macrophage and
displays increased pathogenic potential to humans [2]. The association of bacteria with amoebae, that
are ubiquitous in the environment, protects the bacteria from common disinfection measures. This
leads to the creation of reservoirs of pathogenic microorganisms that are difficult to eradicate [3]. This
process occurs widely in the environment, but is of special concern in hospitals, where many
established and emerging pathogens involved in nosocomial disease, such as Legionella pneumophila,
are increasingly being identified in association with amoebae [4]. One particularly important amoeba-
associated microorganism (AAM) is the newly recognized giant mimivirus [5], a member of the
Nucleo-Cytoplasmic Large Deoxyribovirus (NCLDV) group. The virus has been isolated from
hospital sources in France and, together with other members of the NCLDV group, is known from
metagenomic studies to be widespread in the Oceans, representing one of the main groups of marine
viruses [6]. Our project aims to detect amoeba and potentially pathogenic AAMs with particular
relevance to the NCLDVs in the environment and in hospital facilities. In the present work we have
developed methods for detection, isolation and in vitro culture of amoebae from several environmental
sources including hospital dust, sea water and estuarine sediments.
Suspensions of samples from hospital dust and biofilms and estuarine sediment and water samples
were concentrated by centrifugation at 800rpm and seeded in non-nutrient agar covered with
monolayers of live Lactobacillus spp. or E. coli. Growth of amoeba was detected 3 to 10 days after
seeding, by inspection of the agar plates with a light microscope (Figure 1a). For light microscopy
studies, amoeba suspensions were allowed to attach to glass slides overnight or centrifuged onto the
slides with a cytocentrifuge, fixed in methanol and stained with Giemsa. For transmission electron
micoscopy, the pelleted amoebae were fixed sequentially in glutaraldehyde, osmium tetroxide and
uranyl acetate, followed by dehydration and embedding in Epon-Araldite.
Presently, we are carrying out the characterization of a number of amoebae isolated from the sources
mentioned above. Amoebae have been identified both through morphology, using light and electron
microscopy, and by PCR amplification of rRNA coding sequences and further sequencing of the
obtained amplicons [7]. Most of the isolated amoebae were Acantamoebae with characteristic
morphologic features like the acanthopodia (Figure 1b). Some belong to other genera yet to be
characterized (Figure 1c).
In one sample of amoeba isolated from hospital dust and identified as Acanthamoeba castellanii,
transmission electron microscopy allowed the identification of amoeba-associated virus-like particles
resembling mimiviruses (Figure 1d). Our work continues in order to characterize and isolate amoeba
and AAMs from the studied sources.
31
doi:10.1017/S1431927612012810
Microsc. Microanal. 18 (Suppl 5), 2012
© Microscopy Society of America 2012
https://doi.org/10.1017/S1431927612012810
Downloaded from https://www.cambridge.org/core. IP address: 35.168.9.130, on 16 Feb 2022 at 23:12:44, subject to the Cambridge Core terms of use, available at https://www.cambridge.org/core/terms.