Journal of Microbiology Research 2013, 3(1): 11-18
DOI: 10.5923/j.microbiology.20130301.02
Characterization of Vibrio Isolates from Carpet Shell
Clam (Ruditapes Decussatus ) Suffering from Brown Ring
Disease (BRD) on Tunisian Coasts
Monia El Bour
1,*
, Fatma Lakhal
1
, Radhia Mraouna
1
, Annick Jacq
2
, Christine Paillard
3
, John Klena
4
1
Laboratoire de Pathologie, Institut National des Sciences et Technologies de La Mer (INSTM), 2025, Salammbô, Tunisie
2
Institut de Génétique et Microbiologie, Université Paris-Sud, CNRS, Orsay, France
3
LEMAR UMR, Institut Universitaire Européen de la Mer-CNRS, Plouzané, France
4
U.S. Centers for Disease, AU1 Control and Prevention, Unit 7300, Box 0066, DPO AP, US
Abstract A total of 13 pathogenic Vibrio spp. bacteria were isolated from carpet shell clams (Ruditapes decussatus)
with Brown Ring Disease (BRD) in Tunisia. Organisms were identified based on a combination of phenotypic and
molecular methods (SSP-PCR and 16SrDNA sequencing). Virulence effects were determined by in vivo testing on R.
decussatus and in vitro testing on the Manila clam (Ruditapes philippinarum). All isolates demonstrated biochemical
profiles typical of Vibrio spp; nine different biotypes. Three isolates were identified as V.splendidus biovar II TAE2 but the
remaining isolates fell into eight biotypes different from each other and from the V.tapetis biotype. Species delineation
based on 16S rDNA sequencing indicated that the isolated Vibrio spp closely resembled V.chagasii (eight isolates),
V.splendidus (two isolates), V.alginolyticus(one isolates), Psychrobacter spp. and Pseudoalteromonas mariniglutinosa (one
isolates for each). In vitro cytotoxicity effects and mortalities could be induced by the isolates of V.chagasii, V.splendidus
and V.alginolyticus at lower dosages than induced by V.tapetis (CECT4600).
Keywords Vibrio, Brown Ring Disease, Biochemical Profiles, Sequencing, Toxicity, Ruditapes decussatus, Ruditapes
Philippinarum
1. Introduction
Many bacterial diseases in adult bivalves are caused by
organisms in the genus Vibrio (Gram-negative, rod-shaped
bacteria) that are widespread in coastal and estuarine
environments ([6],[20],[23],[33]). Recent data have shown
an expansion in the host range of aquatic organisms affected
by pathogenic Vibrio species; these are primarily bivalve
invertebrates ([7],[11],[27]). In addition to bivalve
bacteriosis, Brown Ring Disease (BRD), a disruption of the
periostracal lamina with abnormal conchiolin deposition,
which affects main ly the Manila clam Ruditapes
philippinarum and the carpet shell c la m Ruditapes
decussatus. Infections of BRD have been documented in
wild populations and cultured clams isolated from waters
around France, England, Ireland, Spain, occasionally in Italy
and even in Korea ([2],[33],[35],[37]).
In most reports of BRD contaminations, Vibrio tapetis
bacteria have been identified ([2],[10],[30]). V.tapetis – like
* Corresponding author:
monia.elbour@instm.rnrt.tn (Monia El Bour)
Published online at http://journal.sapub.org/microbiology
Copyright © 2013 Scientific & Academic Publishing. All Rights Reserved
organisms, such as those identified in[35], have also been
reported as causing BRD in Manila clams in Korea. BRD
could be induced by exposing stocks of Manila clams to V.
tapetis[13], resulting in high mortality. However, the number
of Vibrio species negatively impacting R. decussatus and R.
philippinarum has increased to more than 12 species,
including Vibrio alginoyticus and the Vibrio splendidus
clade ([7],[8],[9]). Vibrio splendidus and related species
Vibrio celticus, Vibrio tasmaniensis, Vibrio neptunius and
Vibrio furnissii have been characterized as pathogen
inducing mortalities for a number of bivalve species
including R. phillipinarum, Venerupis pullastra, Mya
arenaria and Meretrix meretrix ([5],[23],[39],[40]).
The carpet shell clam represents the most economically
important bivalve in Tunisia and is widely distributed along
the Tunisian shoreline where it is extensively harvested for
local consumption and export to Europe. However, very few
studies have been published documenting the health status of
this species. In 2003 we reported the occurrence of BRD in
several wild populations of carpet shell clam on the Tunisian
coast (Southern Mediterranean Sea) with prevalence rates
ranging between 1 and 40%, but with no mortalities[14].
More recently[17], we described the results of a 6-year
surveillance of BRD in natural populations of carpet shell