© 1995 Nature Publishing Group
is a male's trait value and G
1
is the trait's
genetic variance): the correlation thresh-
old then rises to about 0.25, and it rises
still further, to about 0.5, if the preference
experiences a similar intensity of stabiliz-
ing selection as well. No cycling occurs if
the correlation is below the threshold
value. Instead the population settles into
an equilibrium that may or may not fea-
ture extreme male displays, depending in
part on whether or not exaggerated pref-
erences are produced as a side-effect of
natural selection on female behaviour.
This is an important but unanswered
empirical issue.
So the possibility of evolutionary cycles
hinges on the size of this genetic correla-
tion. How large do we expect it to be in
nature? So far, theorists have been able to
calculate the correlation only under
assumptions that are simplified carica-
tures of the genetics and behaviours that
underlie real traits and preferences.
Predictions for the correlation that are
robust to these poorly known details and
based on measurable quantities would be
useful.
A second wish is for a theory of how the
genetic variation in the preference and
trait changes through time - which of
course requires that we understand the
causes of quantitative genetic variation.
Iwasa and Pomiankowski take these vari-
ances to be constants, which is reasonable
given that the means of the trait and pref-
erence make excursions of only about one
phenotypic standard deviation in their
examples. But that amount of change is
trivial compared to the differences we see
between, say, the male plumages of differ-
ent pheasant specie s. If cycles are involved
in generating such diversity, we will ulti-
mately need a theory for the dynamics of
the variances when means evolve over
many phenotypic standard deviations.
Whether changing variances will inhibit
cycles or amplify them, or perhaps throw
evolution into chaos, is anyone's guess. 0
Mark Kirkpatrick is in the Department of
Zoology, University of Texas, Austin, Texas
78712, USA. Nick Barton is in the Institute
of Cell, Animal, and Population Biology,
University of Edinburgh, West Mains Road,
Edinburgh EH9 3JT, UK.
1. lwasa. P. & Pomiankowski. A. Nature 377. 420-422
(1995).
2. Joyce. J. Rnnegans Wake (Faber, London, 1939).
3. Pomiankowski, A. J. theor. Bioi. 128,
(1987).
4 . Kirkpatri ck, M. in Sexual Selection: Testing the
Alternatives (eds Bradbury, J. W. & Andersson. M. B.)
67-82 (W1Iey. Chichester, 1987).
5. Kirkpatrick, M. & Ryan. M. Natur e 350, 33-38 (1991).
6. Fi sher. R. A. The Genetical Theory of Natural Selection
(Clarendon, Oxford , 1930).
7. Lande. R. Proc. natn. Acad. Sci. U.S.A. 78, 3721- 3725
(1981).
8. Murray, J. D. Mathemati cal Biology, 161- 166 (Springer.
Berl in, 1989).
9 . Ludwig, D., Jones, D. D. & Holling, C. S. J. Anim. Ecol. 47,
315-332 (1978).
10. Turelli, M. Theor. Popular. Bioi. 25, 138-193 (1984).
11. Endler, J. A. Natural Selecti on in the Wild (Princeton Univ.
Press, 1986).
NATURE · VOL 377 · 5 OCTOBER 1995
NEWS AND VIEWS
QUANTUM COMPUTING------------------
Towards an engineering era?
Charles H. Bennett and David P. DiVincenzo
ONE of the paradoxes of quantum theory
is how little its strange foundations
obtrude into everyday life, even while sup"
porting an edifice of chemistry and physics
that explains the existence and properties
of ordinary matter in exquisite detail. To
marvel at the foundations themselves -
in the form, say, of the two-slit experiment
or the Einstein-Podolsky-Rosen effect-
one must generally visit a textbook or a
laboratory. But it is beginning to appear
that the foundations of quantum mechan-
ics will find a fairly direct and visible
application in the new science of informa-
tion processing.
For about ten years, there have been
speculations about the vast and distinctive
powers a quantum computer would have
if one could ever be built. Now it appears
that the subject of quantum computing is
emerging from this visionary prehistory
into a distinct 'experimental' era. Powerful
algorithms exploiting the unique capabili-
ties of a quantum computer are now wide-
ly understood, and systematic searches for
new algorithms are under way. On the
hardware side, the implementation of the
logic gates of a quantum computer is now,
at some rudimentary level, within the
purview of well established experimental
physics. It is still too early to tell when or
whether quantum computing will advance
to a third, 'e ngineering' era of routine
practical application.
At a summer workshop of quantum
computation*, the talks, while still includ-
ing 'visionary era' discussions of Bell's
inequalities, quantum cellular automata
and the 'many-worlds' interpretation, had
moved on to include concrete discussions
of experimental realizations of quantum
logic gates - to the extent that the Uni-
versity of Innsbruck group unabashedly
presented a "realistic model of a quantum
computer" (T. Pellizzari; ref. 1).
Their scheme for implementing quan-
tum logic builds on the details of some of
the hottest trends in atomic spectroscopy,
in particular a technique called 'dark-state
transfer of Zeeman coherence'. A Zee-
man-coherent atom is one in which the
wavefunction is only non-zero for one of
the members of the ground-state multi-
plet (containing Zee man fine structure).
This wavefunction can be moved to the
other member of the multiplet if the atom
is exposed to two optical beams which
couple these two ground-state levels to an
excited state (see figure). Th e rule is first
to turn bea m B
2
on, which leaves the sys-
tem in state 1, but strongly couples states
2 and 3. When B1 is turned on next, the
*Quantum Computation, Institute for Scientific Interchange ,
Turin, Italy, 25 June-7 July 1995.
system is transferred from 1 to the 2-3
complex, but, remarkably, because of
quantum interference the amplitude of
the wavefunction in the excited state 3
always remains very small. Then if B
2
is
turned off, the system wavefunction is
entirely in state 2; turning off B
1
com-
pletes the process.
This kind of spectroscopy has many
advantages as a basic step in quantum
State 1 State 2
In dark-state spectroscopy, a quantum sys-
tem passes from state 1 to state 2 by mixing
with an excited state 3 using coherent radia-
tion (modes 81 and 8
2
).
computation. The instantaneous state is
always 'dark': that is, it cannot suffer
spontaneous emission from the excited
state, as its probability density for being in
this state is vanishingly small (if the beams
are turned on and off slowly enough).
Thus the system always remains in a pure
quantum state. If one of the beams B
2
is a
localized cavity mode in a superposition of
zero- and one-photon states, then the
spectroscopy serves to map that super-
position into the same superposition of
the two Zeeman sublevels. It is this trans-
fer capability which the Innsbruck workers
have extended to solve the hardest prob-
lem of physical quan tum logic gate design:
moving a quantum state undisturbed from
one subsystem (atom) to another.
Several experiments have proved that
different quantum bits (or 'qubits') can be
made to interact controllably, in such a
way that a rudimentary quantum logic
gate is formed. In the work of C. Monroe
and collaborators
2
, it has been successfully
shown that the low-lying states of beryl-
389