pubs.acs.org/JAFC Published on Web 12/31/2009 © 2009 American Chemical Society
1180 J. Agric. Food Chem. 2010, 58, 1180–1188
DOI:10.1021/jf902753f
Relationship of Granule Size Distribution and Amylopectin
Structure with Pasting, Thermal, and Retrogradation
Properties in Wheat Starch
SANDEEP SINGH
Department of Food Science and Technology, Guru Nanak Dev University, Amritsar-143005, India
NARPINDER SINGH*
Department of Food Science and Technology, Guru Nanak Dev University, Amritsar-143005, India
NAOTO ISONO
Graduate School of Bioresources, Mie University, Tsu 514-8507, Japan
TAKAHIRO NODA
National Agricultural Research Center for Hokkaido Region, Memuro, Hokkaido 082-0081, Japan
Starches separated from 18 Indian wheat varieties were evaluated to see relationship of granule
size distribution and amylopectin structure with pasting, thermal, and retrogradation properties.
Average diameter of A-, B-, and C-granules among different starches varied between 23.0 and 28.5,
10.0 and 12.0, and 2.3 and 2.7 μm, respectively. Amylopectin chain length distribution varied
significantly, short length chains (DP 6-12) and long length chains (DP > 24) ranged between 44.5
and 52.4% and 3.7 and 6.5%, respectively, whereas amylose content ranged between 18.2 and
28.8%. Short length chains of amylopectin had inverse relationship with starch gelatinization
temperatures T
o
, T
p
, and T
c
. Starches with higher crystallinity had higher enthalpy of gelatinization
and lower swelling power. Paste characteristics were mainly dependent upon granule type and
all pasting parameters except pasting temperature, showed significant positive correlations with
A-granules and negative with the proportion of B- and C-granule.
KEYWORDS: Granule size; retrogradation; amylopectin; wheat starch
INTRODUCTION
Different size and shapes of starch granules build up in the
endosperm during the development of grain. The mature endo-
sperm in wheat consists of two distinct types of starch granules
that are large, disk-shaped, and termed A-granules as well as
small and spherical, termed as B-granules. Bechtel et al. ( 1 ) and
Raeker et al. ( 2 ) reported for wheat starch having another distinct
class of small C-granules that initiated 21 days after anthesis. The
different granule types show significant differences in chemical
composition and functional properties such as amylose, lipid
content, and gelatinization characteristics ( 3 , 4 ).
Starch is composed of two glucose polymers, amylose and
amylopectin. Amylopectin with its multiple branched chains of
(1-4)-R-glucans interlinked by (1-6)-R-linkages is usually the
major component of starch, with the essentially unbranched amyl-
ose making up the minor fraction. The molecular architecture of
amylopectin and its molecular arrangement within the granule is
related to the granule size ( 5 ). The ratio of long branch chains to
short branch chains affects the shape of the amylopectin mole-
cules, which affects their packing and, in turn, the morphology and
size of the starch granule ( 6 ). The functional properties of starch
are affected by amylose content, branched chain-length distribu-
tion of amylopectin ( 7 , 8 ), phosphate monoester, phospholipid,
and lipids content ( 9 , 10 ), starch granule size distribution ( 2 , 11),
crystalline structures ( 12 ), and granular architecture ( 13 ). In wheat
starches, amylopectin is considered to contribute to water absorp-
tion, swelling, and pasting of starch granules, whereas amylose
and lipids tend to retard these processes ( 14 ). Jane and Chen ( 15 )
concluded that the amylopectin chain length distribution and
amylose molecular size produce synergistic effects on the viscosity
of starch pastes. The present study assesses the characteristics
of starches from different Indian wheat varieties with regard
to structural, physicochemical, granule size distribution and
proportion, pasting, thermal, and retrogradation properties. The
objective of our study was to investigate the structural and
physicochemical features that provide diverse functional pro-
perties to wheat and to develop a relationship between these
properties.
*To whom correspondence should be addressed. Phone: 0183-
2258802 ext 321. Fax: þ91-183-2258820. E-mail: narpinders@
yahoo.com.