ORIGINAL ARTICLE
Enhancement of viability of a probiotic Lactobacillus
strain for poultry during freeze-drying and storage
using the response surface methodology
Anahita KHORAMNIA,
1
Norhani ABDULLAH,
1,2
Siew Ling LIEW,
3
Chin Chin SIEO,
1,2
Kalavathy RAMASAMY
4
and Yin Wan HO
2
1
Department of Microbiology, Faculty of Biotechnology and Biomolecular Sciences,
2
Institute of Bioscience,
Universiti Putra Malaysia,
3
Faculty of Science and Technology, Universiti Kebangsaan Malaysia,
4
Faculty of
Pharmacy, Universiti Teknologi MARA Malaysia, Selangor Darul Ehsan, Malaysia
ABSTRACT
A rotatable central composite design (CCD) was used to study the effect of cryoprotectants (skim milk, sucrose and lactose)
on the survival rate of a probiotic Lactobacillus strain, L. reuteri C10, for poultry, during freeze-drying and storage. Using
response surface methodology, a quadratic polynomial equation was obtained for response value by multiple regression
analyses: Y = 8.59546 - 0.01038 X1 - 0.09382 X2 - 0.07771 X3 - 0.054861 X1
2
- 0.04603X3
2
- 0.10938 X1X2. Based on
the model predicted, sucrose exerted the strongest effect on the survival rate. At various combinations of cryoprotectants,
the viability loss of the cells after freeze-drying was reduced from 1.65 log colony forming units (CFU)/mL to 0.26–0.66 log
CFU/mL. The estimated optimum combination for enhancing the survival rate of L. reuteri C10 was 19.5% skim milk, 1%
sucrose and 9% lactose. Verification experiments confirmed the validity of the predicted model. The storage life of
freeze-dried L. reuteri C10 was markedly improved when cryoprotectants were used. At optimum combination of the
cryoprotectants, the survival rates of freeze-dried L. reuteri C10 stored at 4°C and 30°C for 6 months were 96.4% and
73.8%, respectively. Total viability loss of cells which were not protected by cryoprotectants occurred after 12 and 8 weeks
of storage at 4°C and 30°C, respectively.
Key words: cryoprotectants, freeze-drying, probiotic, response surface methodology, storage.
INTRODUCTION
During the past 50 years, antibiotics have been used in
livestock production at therapeutic levels to treat bac-
terial infections, as well as at sub-therapeutic levels to
promote growth. Many of the antibiotics used as
growth promoters in the livestock industry are also
used in human medicine. Shortly after the initiation of
widespread use of antibiotics in the livestock industry,
particularly the poultry industry, the practice was
placed under increased scrutiny because of concern
over the development of bacterial resistance to the
usual microbiocidal effects of the antibiotics. Since
then, the indiscriminate use of antibiotics in poultry
production has become a cause for concern (Edens
2003).
An understanding of the importance of the intesti-
nal microflora in the maintenance of health and the
prevention of disease has led to the suggestion that
probiotics could be used as an alternative to antibiotic
growth promoters in livestock. Probiotics are defined
as viable microorganisms that when administered in
adequate amounts, promote or support a beneficial
balance of the autochthonous microbial population of
the gastrointestinal tract (Holzapfel & Schillinger 2001;
Fung et al. 2008). As the viability of the probiotic
microorganisms is essential to exert beneficial effects
on the host, the viability and stability of the probiotic
microorganisms during storage are critical factors to be
considered by manufacturers and consumers.
Drying methods are the most common techniques
used for preservation of microorganisms (Carvalho
et al. 2003) and preservation of large quantities of bac-
teria by freeze-drying is frequently used. However, the
Correspondence: Sieo Chin Chin, Department of Microbiol-
ogy, Faculty of Biotechnology and Biomolecular Sciences,
Universiti Putra Malaysia, 43400 UPM Serdang, Selangor
Darul Ehsan, Malaysia. (Email: ccsieo@biotech.upm.edu.my)
Received 12 July 2009; accepted for publication 14 February
2010.
Animal Science Journal (2011) 82, 127–135 doi: 10.1111/j.1740-0929.2010.00804.x
© 2010 The Authors
Journal compilation © 2010 Japanese Society of Animal Science