Volume 105A, number 4,5 PHYSICS LETTERS 15 October 1984
MULTIPLE PERIODIC REGIMES AND FINAL STATE SENSITMTY
IN A BIOCHEMICAL SYSTEM
O. DECROLY and A. GOLDBETER
Faculte des Sciences, Universitd Libre de Bruxelles, C.P. 231, Campus de la Plaine, B-1050 Brussels, Belgium
Received 5 July 1984
We analyze a model biochemical system governed by three nonlinear differential equations, under conditions where
multiple stable periodic regimes coexist. We determine the structure of their basins of attraction and show that t-mal state
sensitivity is obtained when two stable oscillations are separated by unstable chaos. The effect of fractal basin boundaries
on the response of the oscillatory system to perturbations is discussed.
Nonlinear dynamical systems often show the coex-
istence, for the same set of parameter values, of two
or more different attracting states - either stationary
or periodic. The question arises then as to what will
be the time asymptotic regime as a function of initial
conditions. Several authors have investigated this as-
pect of nonlinear dynamics, revealing complex struc-
tures for the basins of attraction of the different
asymptotic regimes [ 1-5]. In addition to simple ba-
sin structure, for which the boundary of the basis of
attraction can be depicted as a simple surface, one
can observe self-similar stripe structure [4], (see also
ref. [2], fig. 9, 10), i.e. a basin constituted by an in-
finity of "stripes" whose sizes decrease exponentially
as they accumulate, and even fractal basin structure
[1-3,5]. In the latter case, as a fine structure for the
limits of the basins is observed at all scales of precision,
it becomes exceedingly difficult to predict the final
state as a function of initial conditions, a situation
which has been referred to as the final state sensiti-
vity [3].
The basin structures have been investigated mainly
on discrete-time, one- [5 ] as well as two-dimensional
[1-3] return maps. As to differential systems, a basin
shape very similar to the stripe structure has been
found in a chemical model for bistability and oscilla-
tions [6]. There is until now, however, no clear de-
monstration of fractal basins in continuous differen-
tial systems, although Grebogi et al. [3] have present-
0.375-9601/84/$ 03.00 © Elsevier Science Publishers B.V.
(North-Holland Physics Publishing Division)
ed evidence for such phenomenon in the Lorenz
model, as well as in experiments in hydrodynamics.
We report here final state sensitivity in a biochemical
model governed by three differential equations, and
determine the consequences of such phenomenon for
the dynamics of the oscillatory system.
The model which represents a sequence of two
autocatalytic enzyme reactions coupled in series (fig.
1) can be viewed as a simple prototype for the inter-
play between two instability-generating mechanisms.
In addition to simple and complex periodic oscilla-
tions, this system may present chaos and birhythmicity
(coexistence of two stable limit cycles under the same
conditions) [7], and even a coexistence between
three stable limit cycles [8]. The time-evolution of
the concentrations of S(a), Pl(/3) and P2(7) is govern-
ed by the following ordinary differential equations,
where the various parameters, besides o, the constant
input, and the sink constant ks, relate to the kinetic
÷ ÷
Fig. 1. Model enzymatic system comprising two autocatalytic
reactions coupled in series. A substrate S is transformed into
a product P1 by an enzyme El, while the second enzyme
E2 converts P1 into product P2. Each enzyme is activated by
its reaction product.
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