Journal of Food and Nutrition Research Vol. 47, 2008, No. 1, pp. 18-22
18 © 2008 VÚP Food Research Institute, Bratislava
Staphylococcus aureus is a Gram-positive coc-
cus, which forms grape-like clusters. This bac-
terium is facultatively anaerobic, mesophilic,
catalase-positive and oxidase-negative. It grows
within a temperature range from 7 °C to 48 °C and
in a pH range from 4.2 to 9.3. Its distinctive fea-
ture is halotolerance equivalent to minimal water
activity (a
w
) values ranging from 0.83 to 0.85 [1].
Strains of these bacteria are characterized by pro-
duction of heat-stable enterotoxins, which may be
produced at a
w
values as low as 0.86 under aerobic
conditions.
S. aureus is generally associated with skin and
mucous membranes of warm-blooded animals, in-
cluding humans. BAIRD-PARKER [2] considered this
organism to be transient and not usually becoming
a part of the resident flora of the skin. Irrespective
of this fact, growth of S. aureus is connected with
artisanal bryndza cheese produced from raw ewes’
milk in Slovakia. The level of milk contamination
by these pathogenic bacteria may be increased
when mastitis is present in the herd. During
milking and primary treatment of milk on farms,
S. aureus may colonize the parts of technological
equipment that are difficult to clean [3]. For exam-
ple, typical counts of S. aureus in properly drawn
ewes’ milk are between 100 and 200 CFU.ml
-1
[4], and S. aureus is considered to be one of the
ubiquitous contaminants. The presence of toxin-
producing strains in ewes’ lump cheese or bryndza
cheese produced from raw milk is dependent on
various factors that include animal health status
and environmental, hygienic and technological
conditions during production. On the other hand,
fermentation and ripening of cheese reduce the
probability of enterotoxin production. According
to [5], increase in the bacterial population during
initial 24 h of cheese production in normal con-
ditions is about 1.5 to 3 log CFU.ml
-1
. The maxi-
mum contents of S. aureus of m = 10
4
CFU.g
-1
and
M = 10
5
CFU.g
-1
, (n = 5, c = 2; where n is number
of units comprising the sample, c – number of
sample units giving values greater than m or be-
tween m and M) is a criterion of process hygiene
for cheeses produced from raw milk according to
Regulation (EC) No. 2073/2005 [6].
When analysing a bacterial growth curve,
lag time and the exponential growth phase are the
most important for growth assessment. At present,
the Baranyi model and Gompertz function are the
most widely used primary models for the descrip-
tion of microbial growth [7, 8].
The growth of a culture of a particular micro-
bial strain in foods is determined by its intrinsic
Variability of growth parameters of Staphylococcus aureus in milk
ĽUBOMÍR VALÍK – ALŽBETA MEDVEĎOVÁ – BARBORA BAJÚSOVÁ – DENISA LIPTÁKOVÁ
Summary
Growth of 15 strains of Staphylococcus aureus in milk at 15 °C was studied in 87 experiments, in order to determine
the reproducibility of the growth parameters measured. The average growth rate (Gr) at 15 °C was 0.067 log CFU.h
-1
,
ranging from 0.54 to 0.77 log CFU.h
-1
, which was equivalent to a doubling time ranging from 12.8 h to 9.0 h. Using
a standard inoculation of an 18 h culture, the initial counts in milk of 3.68 log CFU.ml
-1
and the duration of the lag
phase of 14.4 h were determined with the standard deviations of 0.2 log CFU.ml
-1
and 2.4 h, respectively. Based on
ANOVA tests, the growth rates of individual strains did not produce different variances at the statistically significant
5% level. The results show that despite the errors connected with cultivation methods of determination microbial
counts, the coefficient of variation (CV) of the growth rates for all 15 strains was as low as 7.1%. When 6 growth curves
for a single strain were studied in two independent experiments, a CV of 1.2% for the growth rate was determined.
Keywords
growth parameters; variability; Staphylococcus aureus; predictive microbiology
Ľubomír Valík, Alžbeta Medveďová, Barbora Bajúsová, Denisa Liptáková, Institute of Biochemistry, Nutrition and Health
Protection, Department of Nutrition and Food Safety, Faculty of Chemical and Food Technology, Slovak University of
Technology, Radlinského 9, SK – 812 37 Bratislava, Slovakia.
Correspondence author:
Ľubomír Valík, e-mail: lubomir.valik@stuba.sk