Botanical Journal of the Linnean Society, 2003, 143, 189–196. With 9 figures
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Original Article
AECHMEA MULTIFLORA
COMPLEX
M. B. FERREIRA CANELA
ET AL
.
*Corresponding author. E-mail: twendt@biologia.ufrj.br
Revision of the Aechmea multiflora complex
(Bromeliaceae)
MARIA BERNADETE FERREIRA CANELA, NATÁLIA PRADO LOPEZ PAZ
and TÂNIA WENDT*
Departamento de Botânica, Universidade Federal do Rio de Janeiro, CCS, IB, Ilha do Fundão, 21941–
590, Rio de Janeiro-RJ, Brazil
Received January 2002; accepted for publication May 2003
A taxonomic revision of the Aechmea multiflora complex (Bromeliaceae) is presented. The complex currently com-
prises four species belonging to the subgenus Chevaliera. In this study, we document morphological variation in her-
barium and living plants in order to improve the delimitation of species in the genus Aechmea. Three species were
recognized: A. multiflora, A. depressa, and A. saxicola. We propose A. hostilis to be a synonym of A. saxicola. These
species are restricted to the eastern coast of Brazil, and are known from only a few herbarium collections. The con-
servation status of these species is ‘vulnerable’ according to IUCN criteria. We present a key, descriptions, typifica-
tion, drawings, photographs and a map of distribution. © 2003 The Linnean Society of London, Botanical Journal
of the Linnean Society, 2003, 143, 189-196.
ADDITIONAL KEYWORDS: Brazil - Bromelioideae - Chevaliera - conservation - morphology - taxonomy.
INTRODUCTION
Systematics in the Bromeliaceae is controversial
and problematic. This is particularly true for the
subfamily Bromelioideae in which the genus Aech-
mea Ruiz & Pav. is placed. The most recent mono-
graphic treatment of the Bromelioideae is that of
Smith & Downs (1979), in which they recognized
172 species of Aechmea, dividing them into eight
artificial subgenera. Since 1979, numerous new spe-
cies and generic-level taxonomic changes have been
proposed in the Bromelioideae, many involving the
genus Aechmea (Read, 1984; Smith & Spencer, 1992;
Luther & Sieff, 1994, 1997; Read & Baensch, 1994;
Luther, 2001). The most controversial was the eleva-
tion of all Aechmea subgenera to generic rank
(Smith & Kress, 1989, 1990), an action that has not
been widely adopted (e.g. Luther & Sieff, 1994).
These problems are almost always magnified by an
incomplete knowledge of critical diagnostic mor-
phological features for many taxa. The taxonomic
problems within Aechmea require detailed scrutiny
before the proposal of new treatments (Wendt,
1997).
This paper attempts to provide a sound delimita-
tion of the closely related species which constitute
the A. multiflora complex: A. multiflora L. B. Sm.,
A. depressa L. B. Sm., A. saxicola L. B. Sm., and
A. hostilis E. Pereira. These species belong to the sub-
genus Chevaliera (Gaudich. ex Beer) Baker (sensu
Smith & Downs, 1979) which comprises c. 21 robust
tank bromeliad species, bearing hard serrate leaves
over 1 m long, with large strobiliform inflorescences
commonly armed with pungent bracts. They fre-
quently grow as epiphytes. Due to difficulties in
accessing, collecting, drying, and handling material
from these species, the number of available herbar-
ium specimens is small. Smith & Downs (1979) cite
the type plus five specimens for A. multiflora and
basically the type collections for the other three taxa.
Smith (1937, 1941, 1950) described three species rec-
ognized in this complex but made no comparison or
comments about the affinities between them. Smith
(1937) described A. multiflora, and suggested affini-
ties between the Amazonian A. fernandae (E. Morren)
Baker and Amazonian A. rubiginosa Mez, both of the
subgenus Chevaliera. Later, Smith (1941) described
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