Systematic revision of the Neotropical genus Entomoderes Solier (Coleoptera: Tenebrionidae) GUSTAVO E. FLORES and SERGIO ROIG-JUÑENT The genus Entomoderes(Pimeliinae:Nycteliini) distributedfrom southern Boliviato central Argentina is revised and a cladistic analysis including nine species is conducted. It comprises E. infernalis Burmeister, E. subauratus Burmeister, E. pustulosus Fairmaire, E. satanicus Waterhouse, E. draco Waterhouse, E. borealis Peña, E. erebi (Lacordaire), E. cellulosus Waterhouse, and E. zupay sp. n. The cladistic analysis shows that Entomoderesis a natural group of species characterized by several synapomorphic characters from the external mor- phology and female genitalia. The species relationships are partially resolved. The paper includes a redescription of the genus, redescriptions or descriptions of its nine species, an identification key, drawings of habitus and genitalic features, distribution maps, trees, and a discussionof the phylogeny and distribution of the genus. G. E. Flores and S. Roig-Juñent, Instituto Argentino de Investigaciones de las Zonas Aridas (IADIZA,CRICYT), C.C. 507, 5500 Mendoza, Argentina. Introduction Entomodere.s belongs to Nycteliini, an endemic Neotropical tribe of Pimeliinae, with 285 species that inhabit arid and semiarid lands of Peru, Bolivia, Paraguay, Chile, Argentina, and Uruguay (Flores 1997). The genus comprises nine species that occur from southern Bolivia to central Argen- tina, principally in the biogeographical provinces of Monte and Chaco (Cabrera & Willink 1980). Previous systematic revisions of Entomoderes were exclusively based on external morphology, ie. not including characters of the male and female genitalia (Kulzer 1954; Pena 1980), and indicated a considerable amount of intraspecific variation. The main objectives of this study are to revise Entomoderes by incorporation of new char- acters from external and internal morfology and genital features, and to establish the phylogenetic relationships among its species. Materials and methods The present study is based on the examination of 302 adults specimens, including the type material of some previously described species. The materi- al studied was borrowed from the following insti- tutions and curators (abbreviations identify col- lections in the text): AMNH American Museum of Natural History, New York, USA (Lee Herman) CEUNC Cdtedra de Entomologia, Universidad Nacio- nal de C6rdoba, C6rdoba, Argentina (Miguel A. Delfino) FIML Fundaci6n e Instituto Miguel Lillo, San Miguel de Tucuman, Argentina (Arturo L. Teran) IADIZA Instituto Argentino de Investigaciones de las Zonas Aridas, Mendoza, Argentina (Sergio Roig-Jufient) INESA Instituto de Entomologia de Salta, Rosario de Lerma, Salta, Argentina(Manfredo A. Fritz) MACN Museo Argentino de Ciencias Naturales 'Ber- nardino Rivadavia', Buenos Aires, Argentina (Axel O. Bachmann) MLP Museo de La Plata, La Plata, Argentina (Juan A. Schnack) Body length was measured dorsally, along the midline, from anterior margin of labrum to elytral apex. Main external morphological features con- sidered as well as the name and position of keels and careens are indicated in Fig. 1. Dissection methods are those used by Tschinkel & Doyen (1980) for genital structures, and by Flores (1997)