PII S0031-9384(97)00310-7
The Newborn Rat Ingests Fluids through a
Surrogate Nipple: A New Technique for the
Study of Early Suckling Behavior
EVGENIY S. PETROV, ELENA I. VARLINSKAYA AND
WILLIAM P. SMOTHERMAN
Laboratory of Perinatal Neuroethology, Center for Developmental Psychobiology, Department of Psychology, State
University of New York at Binghamton, Binghamton, NY 13902-6000
Received 15 January 1997; Accepted 14 May 1997
PETROV, E. S., E. I. VARLINSKAYA AND W. P. SMOTHERMAN. Feeding of the newborn rat with a surrogate nipple: A new
technique for the experimental study of early suckling behavior. PHYSIOL. BEHAV. 62(5) 1155–1158, 1997.—In this report we
describe an apparatus and procedure that permits a newborn rat pup to ingest test fluids including milk through a surrogate nipple. The
surrogate nipple represents a new testing situation for the experimental study of sensory and neurochemical controls of suckling
behavior immediately after birth. © 1997 Elsevier Science Inc.
Surrogate nipple Artificial feeding Suckling Rat pup Milk Development
STUDY of the infant rat has served as a focus of psychobio-
logical research for nearly three decades (6) and has provided a
foundation for investigating the development of suckling be-
havior (1,2). Nevertheless, experimental study of the first suck-
ling episode including the physiological and biochemical mech-
anisms that regulate early behavior at the nipple can benefit
from a simple and reliable technique where newborn rat pups
ingest milk and other fluids from a surrogate nipple. With this
technique, psychobiologists, physiologists and nutritionists can
assess initial responses of the newborn rat to stimuli present in
the suckling context, including milk and the surrogate nipple
itself. Experiments can take place under controlled conditions
and be conducted apart from active maternal care. The new
technique will compliment existing strategies for studying early
behavior at the nipple including suckling tests conducted on the
ventral surface of an anesthetized female (1) or presentation of
milk through an oral cannula (5,8).
In this report, we describe the apparatus and procedures
that permit the newborn rat pup to ingest fluids through a
surrogate nipple. The surrogate nipple feeding technique will
provide a unique testing situation for investigating the control
and development of behavior that appears in the context of
suckling and becomes functionally important immediately after
birth.
THE SURROGATE NIPPLE APPARATUS
The surrogate nipple apparatus consists of three main parts: the
surrogate nipple, the fluid reservoir and the glove box (Fig. 1). The
surrogate nipple is constructed from a block of soft vinyl material
(Carolina Biological Supply, Burlington, NC) and fashioned with
a sanding disk attached to a high speed dental drill. The surrogate
nipple is 25 mm long and tapers to a 1-mm diameter at the rounded
tip. The surrogate nipple has a round plate (1.5 mm thick; 5 mm
diameter) made from the same soft vinyl material that is positioned
5 mm from the tip of the nipple. The plate provides a contact point
for the snout when the pup exhibits an oral grasp response of the
surrogate nipple (10) (Smotherman, W. P.; Petrov, E. S.; Varlin-
skaya, E. I. Experimental study of the first suckling episode: Rat
pups ingest fluids through a surrogate nipple. Submitted for pub-
lication). The base of the surrogate nipple is attached to the end of
an angled dental probe to facilitate manual presentation by the
experimenter. A length of PE 10 tubing (0.58 mm inner diameter)
extends through the center and ends flush with the rounded tip of
the nipple.
The PE 10 tubing is connected to a fluid reservoir made from
a 5-mL disposable syringe. The syringe is anchored in a test
tube holder and positioned so as to be level with the head of the
experimental subject. The syringe-tubing-surrogate nipple sys-
tem, when filled with a test fluid, represents an open hydraulic
To whom requests for reprints should be addressed. E-mail: SMOTHER@BINGHAMTON.EDU
Physiology & Behavior, Vol. 62, No. 5, pp. 1155–1158, 1997
© 1997 Elsevier Science Inc. All rights reserved.
Printed in the U.S.A.
0031-9384/97 $17.00 + .00
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