https://doi.org/10.1177/1758155920971823
Avian Biology Research
1–9
© The Author(s) 2020
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DOI: 10.1177/1758155920971823
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Introduction
The Saker Falcon Falco cherrug (hereafter Saker) has a
Palearctic breeding distribution that extends across pre-
dominantly grassland and desert biomes from central
Europe to eastern China.
1
The species has undergone popu-
lation declines over three generations (1993–2012), with a
median estimate of 47% decline (mainly in the former
Soviet states of Central Asia), although uncertainty regard-
ing the scale of this decline has resulted in the species being
categorized as ‘Endangered’ on the IUCN Red List as a
precaution.
2
Nonetheless, large populations exist else-
where in Central Asia, primarily on the Mongolian and
Qinghai-Tibetan Plateaus, which harbour an estimated
62% of the global breeding population.
3
Monitoring to
detect population distribution, size and trends in these
extensive and remote regions is logistically complex,
4
lead-
ing to efforts to identify potential breeding areas using cli-
matic envelope modelling,
5,6
determination of effective
population sizes using genomic approaches,
7
and assess-
ment of population trends by monitoring constant study
plots,
8
or artificial nests,
9
or by inferring mortality rates
from breeding turnover.
10,11
Avian reproductive organs exhibit development changes
with sexual maturation and temporal changes associated
with the stages of the breeding cycle.
12,13
Birds that incubate
eggs develop brood patches, defeathered areas of skin on the
Seasonal variation in gonad physiology
indicates juvenile breeding in the
Saker Falcon (Falco cherrug)
Andrew Dixon,
1,2,3
Janelle Ward,
4
Sarangerel Ichinkhorloo,
5,6
Tuvshinjargal Erdenechimeg,
5
Batbayar Galtbalt,
5,7
Batmunkh
Davaasuren,
5
Batbayar Bold
5,8
and Nyambayar Batbayar
5
Abstract
We describe seasonal changes in the physiology of reproductive organs of Saker Falcons electrocuted on an electricity
power line in Mongolia. Macroscopic examination of the gonads revealed asymmetry in testes size, with bias to the
left body side. This asymmetry declined with seasonal increase in testes size during the main egg-laying period of the
breeding season. In females, 91% possessed both left and right ovaries (N = 161); there was no visible oviduct associated
with the right ovary and it was smaller than the functional left ovary. Both ovaries showed seasonal development in
size, becoming larger during the peak egg-laying period. Hierarchical preovulatory follicular development was recorded
in two females, with ca. 5 mm difference in the diameter of sequential follicles. Both sexes developed brood patches
during the main incubation period, with adults more likely to exhibit brood patches than juveniles. Among juveniles,
at least 82% of females and 91% of males were non-breeders without brood patches. The high electrocution rate at
our studied power line provided a rare opportunity to examine the non-breeding component of the Saker population.
Juveniles predominated in the non-breeding population during the main egg-laying period (89%, N = 65), with the
proportion of adults electrocuted being significantly lower among females. Only a small proportion of juvenile females
exhibited gonadal evidence of breeding, consistent with the low observed frequency of juvenile breeders at nests. The
demographic composition of the non-breeding population is consistent with female mortality rates exceeding that of
males, and potentially indicates incipient population decline.
Keywords
Avian reproductive organs, avian testes, breeding age, ovarian follicles
1
Emirates Falconers’ Club, Abu Dhabi, UAE
2
International Wildlife Consultants Ltd., Carmarthen, UK
3
Reneco International Wildlife Consultants, Abu Dhabi, UAE
4
Wildlife Health Solutions, Papamoa, New Zealand
5
Wildlife Science and Conservation Center of Mongolia, Ulaanbaatar,
Mongolia
6
Mongolian Bird Conservation Center, Ulaanbaatar, Mongolia
7
Centre for Integrative Ecology, School of Life and Environmental
Science, Deakin University, Victoria, Australia
8
Key Laboratory of Animal Ecology and Conservation Biology, Institute
of Zoology, Chinese Academy of Sciences, Beijing, China
Corresponding author:
Andrew Dixon, Reneco International Wildlife Consultants, PO Box
61741, Sky Tower, Al Reem, Abu Dhabi, UAE.
Email: adixonwales@gmail.com
971823AVB 0 0 10.1177/1758155920971823Avian Biology ResearchDixon et al.
research-article 2020
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