Cytotherapy, 2010; 12: 96–106
Correspondence: Patrick C. Baer, Goethe University, Department of Internal Medicine III, Division of Nephrology,Theodor-Stern-Kai 7, 60590 Frankfurt/M,
Germany. E-mail: P.Baer@em.uni-frankfurt.de
(Received 7 August 2009; accepted 28 September 2009)
ISSN 1465–3249 print/ISSN 1477–2566 online © 2010 Informa UK Ltd. (Informa Healthcare, Taylor & Francis AS)
DOI: 10.3109/14653240903377045
Human adipose-derived mesenchymal stem cells in vitro: evaluation
of an optimal expansion medium preserving stemness
PATRICK C. BAER
1
, NADINE GRIESCHE
1
, WERNER LUTTMANN
2
, RALF SCHUBERT
3
,
ARLETTE LUTTMANN
2
AND HELMUT GEIGER
1
1
Division of Nephrology, Department of Internal Medicine III, Goethe University, Frankfurt, Germany,
2
EuroBioSciences
GmbH, Friesoythe, Germany, and
3
Department of Pediatrics, Goethe University, Frankfurt, Germany
Background aims. The potential of cultured adipose-derived stem cells (ASC) in regenerative medicine and new cell thera-
peutic concepts has been shown recently by many investigations. However, while the method of isolation of ASC from
liposuction aspirates depending on plastic adhesion is well established, a standard expansion medium optimally maintain-
ing the undifferentiated state has not been described. Methods. We cultured ASC in five commonly used culture media (two
laboratory-made media and three commercially available media) and compared them with a standard medium. We analyzed
the effects on cell morphology, proliferation, hepatocyte growth factor (HGF) expression, stem cell marker profile and
differentiation potential. Proliferation was measured with a 3-(4,5-dimethylthiazol-2-yl)-2,5-diphenyltetrazolium bromide
(MTT) assay and a fluorescent assay. Release of HGF was assessed by an immunoassay. Expression of characteristic stem
cell-related transcription factors and markers was evaluated by quantitative polymerase chain reaction (qPCR) (Nanog,
Sox-2, Rex-1, nestin and Oct-4) and flow cytometry (CD44, CD73, CD90, CD105 and CD166), and differentiation was
shown by adipogenic medium. Results. The morphology and expansion of ASC were significantly affected by the media
used, whereas none of the media influenced the ASC potential to differentiate into adipocytes. Furthermore, two of the
media induced an increase in expression of transcription factors, an increased secretion of HGF and a decrease in CD105
expression. Conclusions. Culture of ASC in one of these two media before using the cells in cell therapeutic approaches
may have a benefit on their regenerative potential.
Key Words: adipose tissue, culture, differentiation, expansion, medium, mesenchymal stromal cells
Introduction
Adipose-derived mesenchymal stromal cells (ASC)
are multipotent adult mesenchymal stromal cells
(MSC) that have characteristics very similar to
bone marrow (BM)-derived mesenchymal stromal
cells (BMSC) (1–5). Compared with BMSC, ASC
are easier to isolate because cell aspiration does not
require painful procedures or cause donor site injury,
and adipose tissue contains large numbers of stem
cells compared with BM. At present, the cells are
defined by plastic adhesion and their immunophe-
notypic profile, cell morphology and multilineage
differentiation potential (3,6,7). However, cultured
ASC (as MSC in general) seem to be a heteroge-
neous cell population (8) and no marker specifically
and unequivocally identifies undifferentiated ASC.
Several studies have shown that ASC possess the
potential to differentiate into multiple cell types of
mesodermal, endodermal and epidermal origin, i.e.
chondrocytes, osteoblasts, fibroblasts, adipocytes,
smooth muscle cells, neurons, cardiomyocytes, hepa-
tocytes and endothelial cells (2,3,7). We have shown
previously that ASC undergo differentiation towards
the epithelial lineage when treated with all-trans
retinoic acid (ATRA) (9) or conditioned medium
from primary renal tubular epithelial cells (10). This
versatile differentiation potential places ASC at the
forefront of stem cell-based therapies and transplan-
tation, and may therefore become an emerging and
novel organ regeneration tool for the treatment of
damaged tissues (11). Nevertheless, the use of cul-
tured ASC in stem cell-based therapies is impeded by
limitations of proliferation/population doubling and
loss of stem cell properties during ex vivo expansion,
probably because of the culture medium or supple-
ments used. Although non-expanded mesenchymal
stem cells have also been used in cell therapy (12),
the prerequisite to obtain large cell numbers for
treatments is to expand ASC in vitro by maintain-
ing their stem cell nature. Expansion protocols in