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Short Communication
Spray and Pour-On Acaricides Killed Tennessee (United
States) Field-Collected Haemaphysalis longicornis
Nymphs (Acari: Ixodidae) in Laboratory Bioassays
R. A. Butler,
1,3,
J. G. Chandler,
1
K. M. Vail,
1,
C. J. Holderman,
2,
and R. T. Trout Fryxell
1,
1
Department of Entomology and Plant Pathology, The University of Tennessee, Knoxville, TN, USA,
2
Central Life Sciences, Dallas,
TX, USA, and
3
Corresponding author, e-mail: rbutle25@vols.utk.edu
Disclaimer: The content of this publication is solely the responsibility of the authors and does not necessarily represent the offcial
views of the Foundation for Food and Agriculture Research.
Subject Editor: Timothy Lysyk
Received 18 April 2021; Editorial decision 1 June 2021
Abstract
Haemaphysalis longicornis Neumann (Asian longhorned tick) is an exotic and invasive tick species presenting
a health and economic threat to the United States (U.S.) cattle industry due to its ability to transmit pathogens
and infest hosts in large numbers. The objective of this study was to evaluate available products at causing
H. longicornis mortality in a laboratory bioassay. The effcacy of products was evaluated at label rates using
H. longicornis nymphs collected from a cattle farm in eastern Tennessee in two different bioassays (spray or
dip) against untreated controls. After exposure, ticks were transferred to clean petri dishes and checked for mor-
tality at 0, 1, 2, 3, 4, 21, 24, and 48 h post exposure. No mortality occurred in the untreated controls, whereas
all treated ticks were dead within 24 h of exposure (P < 0.0001). These fndings support the hypothesis that
currently available spray and pour-on products are effective at causing H. longicornis mortality. We conclude
that these acaricides can be used as a component to prevent H. longicornis dispersal and for control in the U.S.
Key words: acaricide, management, bioassay, Haemaphysalis longicornis
Haemaphysalis longicornis Neumann (Asian longhorned tick)
(Acari: Ixodidae) is an exotic and invasive pest recently discovered in
the United States (U.S.), with populations now established in several
eastern states (Egizi et al. 2020). This species feeds on a variety of
wildlife, companion animal, livestock, and human hosts (Schappach
et al. 2020; USDA-APHIS, 2020). Recently, this economically im-
portant tick species has been found with Theileria orientalis gen-
otype Ikeda which is associated with diseased cattle (Oakes et al.
2019; Thompson et al. 2020) and newly invasive populations of
this tick can transmit this pathogen (Dinkel et al. 2021). Knowing
the U.S. strain of H. longicornis has a parthenogenetic reproduc-
tive strategy (Hoogstraal et al. 1968; Qiu et al. 2020), it is specu-
lated that the strategy can increase the ticks invasive potential and
cause signifcant blood loss causing death from anemia (Rainey et al.
2018; Oda et al. 2019; Egizi et al. 2020). Current H. longicornis
management strategies are adopted from the ticks’ native range or
include those used for North American tick species (Schappach et al.
2020). In the United States, there are no established or evaluated
H. longicornis management programs and there are limited data on
acaricide effcacy (Schappach et al. 2020).
Previous studies in other countries evaluated systemic and top-
ical products for H. longicornis control. Otaki et al. (2018) found
lotilaner (Credelio, Elanco, Greenfeld, IN), a veterinary drug com-
monly used to treat companion animal ectoparasites, effectively
killed H. longicornis adults when administered to dogs as a chewable
tablet. Similarly, afoxolaner (Nexgard, Merial, Duluth, GA) canine
chewable tablets killed 100% of H. longicornis 48 h after frst expo-
sure (Kondo et al. 2014). Amitraz, commonly formulated as a spray
and cattle pour-on (PO), signifcantly reduced the number of female
H. longicornis on calves (Heath et al. 1980). Topical imidacloprid and
permethrin were 100% effective at killing female H. longicornis ticks
within 4 d after feeding for 3 h on treated dogs (Hagimori et al. 2005).
Journal of Medical Entomology, 58(6), 2021, 2514–2518
doi: 10.1093/jme/tjab115
Advance Access Publication Date: 1 July 2021
Short Communication
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