Biological Journal of the Linnean Society, 2007, 91, 475–482. With 5 figures
© 2007 The Linnean Society of London, Biological Journal of the Linnean Society, 2007, 91, 475–482 475
Blackwell Publishing LtdOxford, UKBIJBiological Journal of the Linnean Society0024-4066© 2007 The Linnean Society of London? 2007
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Original Article
BEHAVIOURAL SYNDROMES IN FIELD CRICKETS
R. KORTET and A. HEDRICK
*Corresponding author. E-mail: rkortet@cc.jyu.fi
A behavioural syndrome in the field cricket Gryllus
integer: intrasexual aggression is correlated with activity
in a novel environment
RAINE KORTET
1,2
* and ANN HEDRICK
3
1
Behaviour, Ecology and Evolution Team (Integrative Ecology Unit), Department of Biological and
Environmental Sciences, University of Helsinki, PO Box 65(Viikinkaari 1), FI-00014 University of
Helsinki, Finland
2
Section of Ecology, Department of Biology, University of Oulu, PO Box 3000, 90014 University of Oulu,
Finland
3
Neurobiology, Physiology and Behaviour, University of California, Davis, One Shields Avenue, Davis,
CA 95616, USA
Received 27 March 2006; accepted for publication 1 September 2006
Behavioural syndromes, or suites of correlated behaviours across different contexts and situations, have recently
drawn attention from evolutionary biologists. In the field cricket Gryllus integer, males are aggressive with one
another and fight vigorously over females and territories. We examined whether aggressiveness with other males
was correlated with activity in a potentially dangerous context (a novel environment) in laboratory-raised virgin
males. Aggressiveness was measured as fighting ability against a weight-matched opponent. First, we measured
each cricket’s latency to become active in a novel environment and latency to emerge from a refuge within a novel
environment. Next, we determined which of two weight-matched males was more aggressive, by pitting the males
together in an agonistic contest and counting the number of fights won by each male. More aggressive males, who
won more fights, had shorter latencies to become active when placed in a novel environment and shorter latencies
to emerge from a safe refuge. These results suggest that a behavioural syndrome exists in G. integer, in which more
aggressive males are also more active in general, and possibly less cautious towards predation risk. © 2007 The
Linnean Society of London, Biological Journal of the Linnean Society, 2007, 91, 475–482.
ADDITIONAL KEYWORDS: antipredator behaviour – boldness – personality – intrasexual dominance.
INTRODUCTION
Recent studies support the existence of behavioural
syndromes, comprising suites of correlated behaviours
across different situations or contexts (Hedrick, 2000;
Gosling, 2001; Sih, Kats & Maurer, 2003; Sih et al.,
2004b; Sih, Bell & Johnson, 2004a; Carere & Eens,
2005). One of the most familiar behavioural syn-
dromes in animals is an aggressiveness/activity syn-
drome, in which individuals behave more or less
aggressively/are more or less active across many con-
texts (Sih et al., 2004a,b). For example, in funnel-web
spiders, some individuals are more aggressive with
conspecifics than others, attack prey more vigorously,
and expose themselves to higher risk of predation
because of their ‘bold’ or incautious behaviour in the
presence of predators (Hedrick & Riechert, 1989,
1990; Riechert & Hedrick, 1993). Aggressiveness
towards conspecifics that is correlated with danger-
ously bold or noncautious behaviour under predation
risk has been demonstrated in a number of species,
including stickleback fish (Huntingford, 1976, 1982;
Bell, 2005) and fishing spiders (Johnson & Sih, 2005).
The existence of behavioural syndromes may offer an
explanation for apparently ‘non-adaptive’ behaviour,
such as sexual cannibalism, high levels of activity
under predation risk, and hyper-aggressiveness that
decreases mating success by startling or driving away
potential mates (Sih et al, 2004a; Bell, 2005; Sih &
Watters, 2005). Moreover, the persistence of behav-
ioural ‘types’ within a species (e.g. ‘bold’ and ‘shy’ indi-
viduals) suggests that there can be more than one
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