Influence of Organic Amendments on Growth and Lead Uptake of Spinach (Spinacia
oleracea L.) Grown in Lead-Contaminated Soil
Abdur Rashid
1,2*
, Tayyaba Naz
1
, Muhammad Mazhar Iqbal
1,3
, Javed Akhtar
1
, Muhammad Saqib
1
, Hafiz Muhammad
Anwar-ul-Haq
1
, Rooh Ullah
2
, Salahudin Kabir
4
, Qiyamud Din Ikram
5
1
Institute of Soil and Environmental Sciences, University of Agriculture Faisalabad, Punjab 38000, Pakistan
2
Ministry of Rural Rehabilitation and Development, Kabul, Afghanistan
3
Soil and Water Testing laboratory Chiniot, Department of Agriculture, Government of Punjab, Lahore, Pakistan
4
Institut Supérieur d'Agriculture de Lille, Lille 59800, France
5
United Nation University, Institute for Environment and Human Security Bonn, Bonn 53113, Germany
Corresponding Author Email: abdurrashiduaf@gmail.com
https://doi.org/10.18280/eesrj.070201 ABSTRACT
Received: 19 February 2020
Accepted: 13 May 2020
Lead (Pb) is a very serious contaminant in soil because of its widespread application in
agricultural, residential and industrial environments and it has ultimately harmful impacts
on plant and human health. Organic amendments such as biochar (BC) and compost may
reduce heavy metal toxicity in a plant. The main focus of the present research work was
to evaluate the synergistic use of BC and compost to enhance the growth of spinach and
to reduce the Pb uptake in spinach (Spinach oleracea L.) grown on Pb-contaminated soil.
The results showed that Pb toxicity at its higher-level (1000 mg Pb kg-1 soil) reduced the
growth, photosynthetic pigments and physiological attributes in 45-days-old spinach.
However, this decrease was less pronounced in BC and compost amended soil. The BC
and compost application enhanced shoot dry weight, total chlorophyll contents,
membrane stability index and relative water contents at both applied rates 500 and 1000
mg kg-1 of Pb in soil. Under low Pb-stress (500 mg Pb kg-1), BC and compost application
increased the shoot dry weight (33.5 and 39.7%), chlorophyll contents (21 and 22.1%),
membrane stability index (18.8 and 17%) and relative water contents (44 and 34.6%).
While at high Pb-stress (1000 mg kg-1) BC and compost increased the shoot dry biomass
(17.8 and 33.3%), chlorophyll contents (31.3 and 32.5%), membrane stability (27.1 and
23.5%) and relative water contents (37 and 32.9%) respectively. Moreover, under low Pb
stress the application of BC and compost reduced the Pb uptake in spinach by 78 and
56%. While under high Pb stress, BC and compost reduced the Pb uptake by 69 and 52%
respectively. Therefore, it can be concluded that organic amendments can ameliorate Pb-
toxic effects in spinach by changing the physiological attributes (total chlorophyll
content, MSI and RWC) under Pb stress. Among the tested amendments, applied BC
proved more effective for reducing Pb toxicity in spinach.
Keywords:
biochar, compost, heavy metal pollution,
immobilization, Spinacia oleracea L
1. INTRODUCTION
Heavy metal pollution of soil is a threatening issue around
the globe. It is not only responsible to affect the quality and
yield of crops, human and animal health but also the
environment [1]. The major sources of heavy metal pollution
in soil can be find in sewage sludge, industrial waste, mining,
excessive pesticides and chemical fertilizers use and smelting
of ores [2]. These sources have caused a recent gradual
increase in heavy metal concentration and are swept over the
self-purification ability of nature [3]. Subsequently, these
heavy metals persist for a long period, which can be up-taken
by plants to accumulate in plant tissues and hence enter to food
chain [4]. These trace elements are entering to living bodies
via different means like air, food, water and skin contact.
Major health problems linked with heavy metals are mild
cognitive impairment (MCI), chronic anemia and
cardiovascular diseases [5], damage of nervous system, kidney,
brain [6], bones, teeth, skin [7]). Some of these trace elements
are carcinogenic, endocrine disruptors, teratogenic and
mutagenic whereas, others can cause behavioral and
neurological disorders especially in children [8]. These trace
elements at a high concentration are obstructing both growths
of underground and above surface parts of plants which can
cause to inhibit the development and growth of plants and also
to disturb many physiological and biochemical activities e.g.
reduction in transpiration rate, damage the photosynthetic
apparatus, cause breakdown synthesis, injury of the cell
membrane, increase lipid peroxidation and also affect several
enzymes [9].
Among heavy metals, Pb is considering one of the most
hazardous toxic elements and thus it is biologically non-
essential in nature. Lead occurs extensively in the environment
due to its wide spread applications in petrol, paints, explosive
materials, industrial wastes, and sludge. Uptake and
translocation of Pb occur in plants which causes toxic impacts
on its health and ultimately a declination occurs in the plant
biomass production. Usually, plant avoids its toxic effects by
adopting different cellular mechanisms such as adsorption to
the cell wall, vacuole compartmentalization, increase the
Environmental and Earth Sciences Research Journal
Vol. 7, No. 2, June, 2020, pp. 53-61
Journal homepage: http://iieta.org/journals/eesrj
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