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The Pharma Innovation Journal 2019; 8(1): 19-22
ISSN (E): 2277- 7695
ISSN (P): 2349-8242
NAAS Rating: 5.03
TPI 2019; 8(1): 19-22
© 2019 TPI
www.thepharmajournal.com
Received: 13-11-2018
Accepted: 15-12-2018
Shilpi Rajput
Department of Botany,
University of Rajasthan, Jaipur,
Rajasthan, India
Payal Lodha
Department of Botany,
University of Rajasthan, Jaipur,
Rajasthan, India
Correspondence
Shilpi Rajput
Department of Botany,
University of Rajasthan, Jaipur,
Rajasthan, India
Histochemical localization of lipids, and lignin in
healthy and Meloidogyne incognita, infected sunflower
(Helianthus annuus Linn.)
Shilpi Rajput and Payal Lodha
Abstract
The root knot nematodes, belonging to the genus Meloidogyne, which have a wide host range, are the
deadliest enemy of vegetable crops and a pest of crucial economic importance. These produce
conspicuous galls on the roots and the infestations can be recognized easily in fields. It is very common
and abundant in the sandy soils of Jaipur and adjoining areas. In Jaipur district, Sunflower an
economically important vegetable crop was found infested with highly pathogenic Meloidogyne
incognita. Histochemical tests were applied to localize lipids and lignins in the diseased and healthy root
tissues. The metabolites viz. lipids and lignins were found more in galled roots as compared to healthy
roots.
Keywords: Meloidogyne incognita, histochemical localization, root knot nematode, sunflower, metabolites
Introduction
Human kind depends in a myriad ways on plant and plant products. The quest for new plant
products and new methods of using them to satisfy the emerging needs is an ever expanding
enterprise. Unfortunately, these plants become the victim of many disease causing micro-
organisms including the plant parasitic nematodes which can bring about disruption in the
physiological equilibrium of the attacked plants. The plants, in turn, react in a number of ways
to off-set these disturbances and the overall plant response determines, to a large extent, the
success or failure of the interactions. For a better understanding of various histopathological
changes that occurred as a result of nematode infection, in situ localization of various
metabolites was helpful. Although, some pathogens used mechanical force to penetrate plant
tissues but subsequent development of disease syndrome was dependent on histochemical
alterations and biochemical reactions taking place between substances secreted by the
pathogens and those already present or produced by the host as a response to the infection.
Histochemical techniques were advantageous as they enabled in situ localization of various
metabolites at the site of their synthesis or action. While information is available on
morphological and biochemical changes that occur in plants invaded by endoparasitic
nematodes, little work has been done with a histochemical technique. Since the anatomical
area that is biochemically affected by such nematode infections may be quite small, it is
imperative that histochemical techniques be employed, lest the effects of infection be lost by
dilution from non-affected cells.
After histochemically studying the soybean roots infected by Meloidogyne sp., it was reported
that giant cell walls contained cellulose and pectin but lacked lignin, suberin, starch or
ninhydrin positive substances (Dropkin and Nelson, 1960)
[9]
. It was observed that M. javanica
induced giant cell in tomato contained traces of carbohydrates and fats but it was particularly
rich in protein and RNA. The large irregularly shaped nuclei contained a large nucleolus and a
number of feulgen-positive bodies, scattered irregularly along the nuclear envelope (Bird,
1961)
[2]
.
It was reported that in ginger infected with root-knot nematode, giant cell nuclei and
cytoplasm were rich in nucleic acids. Starch was absent in the giant cells and in the cells of the
infected region of the rhizome.
Giant cells showed the presence of minute protein granules. The outer side of the egg sac
consisted of a thick layer of insoluble polysaccharides (Shah and Raju, 1977)
[20]
.