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Vector-Borne Diseases, Surveillance, Prevention
Abundance of Ticks (Acari: Ixodidae) and Presence of
Rickettsia and Anaplasma in Ticks Infesting Domestic
Animals From Northern India
Brij Ranjan Misra,
1
Niraj Kumar,
1
Rajni Kant,
1
Hirawati Deval,
1
Rajeev Singh,
1
Ashok Kumar Pandey,
1
Sthita Pragnya Behera,
1
and Vijay P. Bondre
2,3,
1
ICMR-Regional Medical Research Centre Gorakhpur, Gorakhpur, Uttar Pradesh, India,
2
ICMR-National Institute of Virology, Pune,
Maharashtra, India, and
3
Corresponding author, e-mail: vpbondre@gmail.com
Subject Editor: Kevin Macaluso
Received 15 August 2020; Editorial decision 13 December 2020
Abstract
Rickettsia and Anaplasma are bacteria that can be transmitted by hematophagous arthropods such as ticks
infesting animals in close proximity to humans. The main objective of the present study was to investigate
abundance of common tick species infesting domestic animals and presence of Rickettsia and Anaplasma
in tick populations. Adult ticks were collected from domestic animals in rural areas and screened by molec-
ular detection of bacterial DNA for these two genera of bacteria. A total of 1,778 adult ixodid tick specimens
were collected from 200 cattle, 200 buffaloes, 200 goats, and 40 dogs. The collection consisted of four species
of ixodid ticks, Rhipicephalus (Boophilus) microplus (Canestrini) (83.8%), Hyalomma kumari (Sharif) (7.1%),
Rhipicephalus sanguineus (Latreille) (6.4%), and Dermacentor auratus (Supino) (2.7%) infesting the domestic
animals. The prevalence of all the collected tick species was highest in the month of October. Anaplasma spp.
was the most frequently identifed bacteria (3.3%) in tested ticks. Of 17 positive tick pools for Anaplasma spp.,
14 pools were from ticks infesting cattle, 2 pools of ticks collected from buffalo, and the remaining pool were
ticks infesting a goat at the time of collection. Although 1.6% tick pools of R. microplus collected from cattle
tested positive for Rickettsia spp., present investigation provides evidence of the most prevalent ixodid ticks
infesting domestic animals and the presence of obligate intracellular bacteria, Rickettsia and Anaplasma, in
these ticks collected in the Gorakhpur division of Northern India.
Key words: ixodid tick, PCR, Rickettsia spp., Anaplasma spp.
Ticks are considered one of the most important vectors of Gram-
negative obligate intracellular alphaproteobacteria (Brites-Neto
et al. 2015). Anaplasma and Rickettsia are members of this group
of bacteria. Rickettsia infections are well established as a zoonosis in
various parts of East Asia and India (Ghosh and Nagar 2014). Tick-
borne rickettsial diseases and anaplasmosis are the major illnesses
associated with managing livestock health worldwide (Jongejan and
Uilenberg 2004). Ticks are the major arthropod vectors that transmit
these pathogens from their animal hosts to humans and can also act
as reservoirs in nature (Brites-Neto et al. 2015).
Animals or humans are exposed to ticks and the pathogens that
are harbored by them when they enter into well-established natural
focus (biocenose) where tick-borne pathogen is naturally circulated
between vertebrate host and its tick vector. This situation prevails
more frequently with increasing human population and as more
human activity may occur in the tick-infested habitats (Geevarghese
and Mishra 2011). Usually, ticks have large host range, which
facilitates the shift from one animal host to other hosts which are
in abundance in its habitat. This means ticks parasitizing domestic
animals in human proximity have more chances of accidental con-
tact and frequent bites to humans (Léger et al. 2013). Therefore,
host preference and selection can play a crucial role in spread of
tick-borne bacteria. Hence, study of distribution of ticks and bac-
teria harbored by them is necessary for designing of appropriate in-
tervention strategies.
Seroprevalence for spotted fever (36.7%) and typhus group
(15.3%) of Rickettsia was detected among acute febrile illness cases
during monsoon period (rainy season) from Gorakhpur, India (Mane
et al. 2019). However, no entomological investigations were per-
formed to understand the common tick species and circulation of
Rickettsia and other related bacteria in this area. Therefore, it was
prudent to study distribution of ticks infesting the domestic animals
and presence of Rickettsia spp. and Anaplasma spp. in ticks from
rural areas of Gorakhpur, India.
Journal of Medical Entomology, 58(3), 2021, 1370–1375
doi: 10.1093/jme/tjaa296
Advance Access Publication Date: 20 January 2021
Research
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