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COMMUNICATION
Self-Assembled Autophagy-Inducing Polymeric
Nanoparticles for Breast Cancer Interference In-Vivo
Yi Wang, Yao-Xin Lin, Zeng-Ying Qiao, Hong-Wei An, Sheng-Lin Qiao, Lei Wang,
R P Yeshan J Rajapaksha, and Hao Wang*
Y. Wang, Y.-X. Lin, Dr. Z.-Y. Qiao, H.-W. An, S.-L. Qiao,
Prof. L. Wang, R. P. Y. J. Rajapaksha, Prof. H. Wang
CAS Key Laboratory for Biological Effects of
Nanomaterials and Nanosafety
National Center for Nanoscience
and Technology (NCNST)
No. 11 Beiyitiao, Zhongguancun 100190
Beijing, PR China
E-mail: wanghao@nanoctr.cn
Y. Wang, Y.-X. Lin, H.-W. An, S.-L. Qiao
University of Chinese Academy of Science (UCAS)
No.19A Yuquan Road, 100049 Beijing, PR China
DOI: 10.1002/adma.201405926
one-pot synthesis method and effectively delivered drugs.
[19,20]
Although numerous nanomaterials influence the autophagy
process,
[21]
few examples, to the best of our knowledge, have
been reported to effectively control autophagic flux and ulti-
mately achieve therapeutic potential.
Herein, we describe the development of autophagy-inducing
peptides engineered into polymeric nanoparticles for highly
efficient induction of autophagy and interference of breast
cancer in vitro and in vivo. Bec1, as an autophagy-inducing pep-
tide, covalently grafts onto pH-sensitive polymers (Ps), which
self-assemble into micelle-like nanoparticles (P-Bec1) with
poly(ethylene glycol) (PEG) as hydrophilic shells ( Scheme 1).
The P-Bec1 nanoparticles improved the stability and enhanced
the cellular uptake of Bec1 in vitro. P-Bec1 can dissociate in the
weakly acidic lysosomal environment (pH 4–5), which resulted
in the alkalization and impairment of lysosomes.
[22]
The disso-
ciated P-Bec1 can escape from lysosomes and effectively induce
autophagy, leading to autophagic cell death. Moreover, P-Bec1
is also able to effectively inhibit the growth of tumors and slow
down tumor development in vivo.
The amphiphilic poly(β-amino ester) copolymer composed
of the hydrophobic monomer 1,6-hexanediol diacrylate (HDDA,
1.2 mM), the pH-sensitive monomer 3-(dibutylamino)-1-pro-
pylamine (DBPA, 0.7 mM), and hydrophilic amino-terminated
PEG (PEG-NH
2
, M
W
= 2 kDa, 0.3 mM) was synthesized by
Michael addition.
[19,20]
The structure of the polymer was charac-
terized by
1
H NMR (Figure S4, in the Supporting Information,
SI). The degree of polymerization (DP) and molecular weight
( M
W
) of copolymers were deduced to be 27 and ca. 16 kDa,
respectively.
Furthermore, Bec1 was engineered to have a thiol group
at the N terminal to conjugate with the poly(β-amino ester)
copolymer (Figures S2–S4, SI). The
1
H NMR spectrum of
P-Bec1 showed the disappearance of the acrylate double bonds
of the polymer P at 5.8–6.3 ppm and the appearance of Bec1
peaks at 6.7–8.5 ppm ( Figure 1a), indicating that the acrylate
groups on both ends of the polymer had completely reacted
with Bec1. Circular dichroism (CD) spectra of Bec1 and P-Bec1
were analyzed using standard CONTINLL algorithms,
[23]
which
revealed the secondary structure of Bec1 to be 33.8% α-helix,
and P-Bec1, 18.8% α-helix, proving that 56% of the secondary
structure of Bec1 was not destroyed during the process of copol-
ymer synthesis (Figure 1c). The amphiphilic P-Bec1 polymers
simultaneously self-assemble into micelle-like nanoparticles
during dialysis from dimethyl sulfoxide (DMSO) into water
(Scheme 1). The morphologies and hydrodynamic sizes of P
and P-Bec1 were observed by transmission electron microscopy
Autophagy is a lysosome-based evolutionarily conserved pro-
cess that plays an important role in cellular degradation for the
clearance of damaged or superfluous proteins and organelles.
[1]
Abnormalities in autophagy may directly correlate to various
pathologic diseases, including neurodegenerative disease,
[2]
cardiac disease,
[3]
and cancer.
[4]
Recently, researchers have
attempted to regulate autophagy and prevent further tumor pro-
gress.
[5,6]
Compared to present cancer therapy strategies, such
as chemo-,
[7]
immuno-,
[8]
gene,
[9]
radiation,
[10]
photothermal,
[11]
and photodynamic
[12]
therapies, the autophagy-based concep-
tual pilot studies for cancer therapy are of particular – and
growing – interest in autophagy-deficient cancers, for example,
breast, ovarian, and prostate cancers.
[6]
Beclin-1 (Bec1) is the peptide encoded by the human gene
BECN1, a mammalian homolog of the yeast autophagy-
related gene (Atg) 6, and participates in the regulation of
autophagy through binding to phosphatidylinositol-4,5-bispho-
sphate 3-kinases (PI3Ks, signal transducer enzymes), which
are required for initiation of autophagosome formation in
autophagy.
[13]
BECN1 has been identified as a haploinsufficient
tumor suppressor, and is commonly deleted in human breast,
ovarian, and prostate tumors.
[14]
Furthermore, upregulation of
Bec1 enhances Ras-induced cell death in MCF-7 breast cancer
cells.
[15]
Therefore, it has been suggested that overexpression of
Bec1 could inhibit tumor development.
[6,16]
However, the thera-
peutic Bec1 peptide has two major drawbacks that hamper their
in vivo application, namely poor chemical stability in vivo and
non-specific bio-distribution in tissues. Chemical modification
and/or self-assembly are two useful approaches for overcoming
these limitations.
[17]
pH-sensitive polymers have been devel-
oped widely to realize the controlled disassembly and release of
payloads in biological conditions.
[18]
Our previous studies dem-
onstrated that a pH-sensitive poly(β-amino ester) copolymer
self-assembled into a micelle-like nanoparticle by means of a
Adv. Mater. 2015, 27, 2627–2634
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