J Zool Syst Evol Res. 2020;00:1–10. wileyonlinelibrary.com/journal/jzs | 1 © 2020 Blackwell Verlag GmbH
1 | INTRODUCTION
Our knowledge of the global fauna is continuously enriched with
descriptions of new taxa. Over the past 10 years, the species list
of the family Glossiphoniidae (flat leeches) increased from 208
(Sket & Trontelj, 2008) to 220 (Beresic-Perrins, Govedich, Banister,
& Shuster, 2017; Fedorova & Kaygorodova, 2016; Grosser, Pešić,
Berlajolli, & Gligorović, 2016; Iwama, Nogueira, & Gonsalves, 2017;
Received: 2 July 2019
|
Revised: 3 March 2020
|
Accepted: 12 March 2020
DOI: 10.1111/jzs.12385
SHORT COMMUNICATION
Species delimitation through DNA barcoding of freshwater
leeches of the Glossiphonia genus (Hirudinea: Glossiphoniidae)
from Eastern Siberia, Russia
Irina Kaygorodova
1
| Nadezhda Bolbat
1,2
| Alexander Bolbat
1
Contributing authors: Nadezhda Bolbat (mandzyak@lin.irk.ru), Alexander Bolbat
(bolbatav@lin.irk.ru).
1
Limnological Institute of Siberian Branch of
Russian Academy of Sciences, Irkutsk, Russia
2
Department of Plant Physiology, Cell
Biology and Genetics, Irkutsk State
University, Irkutsk, Russia
Correspondence
Irina Kaygorodova, Limnological Institute
of Siberian Branch of Russian Academy of
Sciences, 3 Ulan-Batorskaya st., Irkutsk
664033, Russia.
Email: irina@lin.irk.ru
Funding information
Russian state assignment, Grant/
Award Number: 0345-2019-0002
(AAAA-A20-116122110066-1); Russian
Foundation for Basic Research, Grant/Award
Number: 17-29-05097
Abstract
The study of biodiversity is a priority task of biological science. The structural unit
of biodiversity is a species that has a clear identification in a taxonomic system.
Morphological features are traditionally the main criteria for species discrimination
in zoological studies. However, the presence of inter- and intraspecific polymorphism
and phenotypic plasticity makes it difficult to identify species in many groups of in-
vertebrates. To solve this problem, in this research, we analyzed morphological and
genetic data in combination to delimit species among the Eastern Siberia Glossiphonia
leeches using different approaches. Morphology analysis revealed phenetically dis-
tinct groups, suggesting the existence of at least two species in the region, G. ver-
rucata, a rare Palaearctic species, and a potentially new species Glossiphonia sp.
Moreover, sequence-based species delimitation methods congruently supported
eight distinct species groups (including two Siberian species) within the available
molecular dataset of the Glossiphonia world fauna, using phylogenetic (ML and BI),
coalescent (ABGD and GMYC) methods, and pairwise analysis of sequences. The de-
tected p-distances (modal value of 0.11) between these 8 groups and the level of
genetic polymorphism (max. 0.0041) within groups indicate that the groups are 8
independent species according to the DNA barcoding. Our results once again proved
the usefulness of molecular systematics. At the same time, we detected several inac-
curacies in the leech species identification, as well as many ambiguous sites in se-
quences uploaded on GenBank, which affects the analysis and impedes progress of
DNA barcoding technology.
KEYWORDS
Eastern Siberia, genetic distances, Glossiphonia, molecular taxonomy, species delimitation