Botanical ]uumal uj the Unllfan Socif!r• 12000!, 134: 19-.19. With 113 figures
doi: 10.1 006/bojl.2000.0:163, availabk at http:/ /www.idealibrary.com on
Under the rnicroscope: plant anatorny and systernatics.
Edited b_y P. ]. Rudall and P. Casson
Pollen morphology and wood anatomy of the
Crudia group (Leguminosae, Caesalpinioideae,
Detarieae)
HANNAH BANKS AND PETER GASSON* FLS
Royal Botanic Gardens, flew, Richmond, Sumy, TW9 3AB
Pollen from all 12 and wood from II genera in the Crudia group haw been examined using
light, scanning electron and transmission electron microscopy. This group is currently of
systematic interest because it is part of a tribe undergoing taxonomic revision. The pollen
grains fall into four groups: (I) Oxystigma, h'ingiodendron, Gossu·eilerodendron, Bathiaea,, Veoapalo>;rlon,
Stemonocoleu.1, Guibourtia and Prioria have pollen of a widespread and generalized cacsalpinioid
type that arc small to medium sized, spheroidal to prolate, tricolporate and with a perforate
exine, with some variation in surface ornamentation, aperture margins and ultrastructure.
(2) Crudia pollen is tricolporate. coarsely striate with a coarsely scabratc to vcrmiculate
aperture membrane. (3) Augouardia is tricolporate and coarsely reticulate. (4) Hardu·ickia and
Colophosperrnum are pantoporate and reticulate or microreticulate-rugulate. The wood of
Prioria, O>;r.1tigma, Kingiodendron and Gos.1weilerodendron has diffusely arranged axial canals, and
these arc four genera that have recently been merged into Prioria. Bathiaea has tangt>ntially
arranged axial canals. The other genera lack normal axial canals. Crudia is distinct, with
banded parenchyma and variably storied short rays, Augouardia has much less abundant axial
parenchyma that is mainly scanty paratrachcal and vasicentric, Guibourtia has mainly aliform
parenchyma and rays variable in height and width, and Colopho.1permum and Hardwickia have
similar paratracheal parenchyma patterns, although the rays tend to be wider in the latter.
Our conclusion is that the Crudia group is not monophyletic.
© 2000 The Linnea I\ Sorict·• of London
ADDITIO:\'AL KEY \\'ORDS:-Augouardia - Bathiaea Colopho,,pmnum Gossznilerodendrorz
- Guibourtia - Hardwickia -- Kirzgiodendron - .Veoapalo>;rlon - O>;rstir;ma - Prioria - Stemonocoleu.l.
Introduction .
Material and methods
Pollen morphology .
Key to pollen .
O:tystigma, Bathiaea, J'v'eoapalo:qlon
Prioria, ( ;osm-eilerodendron
Kingiodendmn
Stemonoculeu.,
Guibourtia
Crudia
Augouardia
CONTE:\'TS
*Corresponding author. E-mail: p.gasson@rbgkew.org.uk
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© :ZOOO The Linne an Socict\· of' London
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