1 Resistant tomato restricts colonization and invasion by the pathogen Ralstonia 1 solanacearum at four organismal levels 2 3 Marc Planas-Marquès 1,± , Jonathan P. Kressin 2,3a,¥,± , Anurag Kashyap 1 , Dilip R. 4 Panthee 3a , Frank J. Louws 2,3b , Nuria S. Coll 1,*, , Marc Valls 1,4,*, 5 6 1 Centre for Research in Agricultural Genomics (CRAG), CSIC-IRTA-UAB-UB, Campus 7 UAB, Bellaterra, 08193 Barcelona, Catalonia, Spain. 8 2 Department of Entomology and Plant Pathology, North Carolina State University, 9 Raleigh, NC 27695, USA. 10 3 Department of Horticultural Science, North Carolina State University, a Mountain 11 Horticultural Crops Research and Extension Center, Mills River, NC 28759, USA and 12 b Raleigh, NC, 27695. 13 4 Department of Genetics, University of Barcelona, 08028 Barcelona, Catalonia, Spain. 14 ± MPM and JPK should be considered joint first authors. 15 * NSC and MV should be considered joint senior authors. 16 ¥ Current address: Department of Breeding, Hortigenetics Research (S.E.Asia) Ltd., 17 East-West Seed Co., Chiang Mai, Thailand 50290 18 19 To whom correspondence should be addressed: Nuria S. Coll 20 (nuria.sanchezcoll@cragenomica.es) and Marc Valls (marcvalls@ub.edu) . Phone 21 number: +34 5636600 22 23 E-mail addresses: 24 Marc Planas-Marquès: marc.planas@cragenomica.es 25 Jonathan P. Kressin: jpkressi@gmail.com 26 Anurag Kashyap: anurag.kashyap@cragenomica.es 27 Dilip R. Panthee: dilip_panthee@ncsu.edu 28 Frank J. Louws: frank_louws@ncsu.edu 29 Nuria S. Coll: nuria.sanchezcoll@cragenomica.es 30 Marc Valls: marcvalls@ub.edu 31