This article has been accepted for publication and undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process, which may lead to differences between this version and the Version of Record. Please cite this article as doi: 10.1111/jse.12695. This article is protected by copyright. All rights reserved. Accepted Article Zhi-yong Zhang ORCID iD: 0000-0003-0490-4181 Research Article Phylogeny and biogeography of Fagus (Fagaceae) based on 28 nuclear single/low-copy loci Lu Jiang 1,2 , Qin Bao 1 , Wei He 1 , Deng-Mei Fan 1 , Shan-Mei Cheng 1 , Jordi López-Pujol 3 , Myong Gi Chung 4 , Shota Sakaguchi 5 , Arturo Sánchez González 6 , Aysun Gedik 7 , De-Zhu Li 8 , Yi-Xuan Kou 2 *, and Zhi-Yong Zhang 2 * 1 College of Forestry, Jiangxi Agricultural University, Nanchang 330045, China 2 Laboratory of Subtropical Biodiversity, Jiangxi Agricultural University, Nanchang 330045, China 3 Botanic Institute of Barcelona (IBB, CSIC-Ajuntament de Barcelona), Passeig del Migdia s/n, 08038 Barcelona, Catalonia, Spain 4 Department of Biology and the Research Institute of Natural Science, Gyeongsang National University, Jinju 660-701, Republic of Korea