Vol.:(0123456789) 1 3
Plant Biotechnology Reports
https://doi.org/10.1007/s11816-020-00598-6
REVIEW
Genetic engineering approaches to understanding drought tolerance
in plants
Zabta Khan Shinwari
1
· Sohail Ahmad Jan
2
· Kazuo Nakashima
3
· Kazuko Yamaguchi‑Shinozaki
4
Received: 20 November 2019 / Accepted: 21 January 2020
© Korean Society for Plant Biotechnology 2020
Abstract
Abiotic stresses such as drought, salinity, frost, etc., afect plant yield manyfold. These stresses can decrease the plant yield of
important major crops up to 50%. The abiotic stress-related genes or other transcription factors (TFs) have multiple functions,
as it increases proline content, leads closing of stomata to decrease the transpiration rate, enhances the production of some
important stress-related protective enzymes, etc. and hence increases abiotic stress tolerance. Many TFs and other stress-
related genes have been identifed and characterized and transformed to many important cultivated plants against drought
and others abiotic stresses. The transformed plants show better morpho-biochemical and physiological performances than
non-transgenic plants. Many genetically engineered plants have been developed against drought stress including wheat, rice,
tomato, soybean, cotton and many more. The efciently engineered clustered regulatory interspaced short palindromic repeats
(CRISPR)/CRISPR-associated nuclease 9 (Cas9) system is now becoming a preferred choice of researchers to edit plant
genomes for introgression natural resistance against a range of abiotic stresses. It leads genome editing by precise manure
with minimal or no efect on growth and development of plants. Very limited reports are available to develop drought-tolerant
plants using CRISPR/Cas9 system. Here we discuss transgenic plant technology and new [CRISPR Cas9 and Virus-Induced
Gene Silencing (VIGS)] techniques to confer drought tolerance in important plant species.
Keywords Abiotic stress · CRISPR/Cas9 · Drought stress · Transcription factor · Transgenic plants · VIGS
Introduction
Global food security is one of the key issues due to change
of world climate condition and by the increase of population.
Drought, salinity, heat, etc., stresses signifcantly afect
plant yield and productivity (Nouri et al. 2015; Singh
et al. 2018). Drought and salinity afect about one-third
of our cultivated land and it cause loss of approximately
1,500,000 ha crop land/year (Peng et al. 2011). In recent
years, drought stress signifcantly decreased the plant yield
manyfold by disturbing its morpho-biochemical processes
(Azevedo et al. 2011). Even lower heat and drought stress
afect plant yield. These stresses reduced crop yield by up to
50%. The plants in reproductive stages are more sensitive to
these stresses and thus afect yield of many important plant
species (Lamaou et al. 2018).
Plants are more susceptible to these extreme
environmental stresses as compared to any other living
organisms. Plants respond poorly to high environmental
stresses, as it afects both biochemical and physiological
processes. So, the development of new engineered plants
is important to fght against these stresses (Ramonell and
Somerville et al. 2002). Decreasing the water amount up
to 40% declines the yield of maize and wheat by as much
as 40% and 21% (Daryanto et al. 2016). The important
cultivated cowpea yield is afected up to 68% by drought
stress (Farooq et al. 2017). About 40% of soybean yield loss
occurs due to drought stress (Specht et al. 1999). It also
reduces cell division, spreading of leaf surface, retards stem
growth and root propagation (Anjum et al. 2015). Long-term
Online ISSN 1863-5474
Print ISSN 1863-5466
* Zabta Khan Shinwari
Shinwari2008@gmail.com
* Sohail Ahmad Jan
sjan.parc@gmail.com
1
Pakistan Academy of Sciences, Islamabad, Pakistan
2
Department of Biotechnology, Hazara University Mansehra,
Mansehra, Khyber Pakhtunkhwa, Pakistan
3
Japan International Research Center for Agricultural
Sciences, Tsukuba, Ibaraki, Japan
4
The University of Tokyo, Tokyo, Japan