A MODEL FOR DIFFUSION AND COMPETITION
IN CANCER GROWTH AND METASTASIS
G.P. PESCARMONA*, M. SCALERANDI**, P.P. DELSANTO**, C.A. CONDAT***
*Dipartimento di Genetica, Biologia e Chimica Medica, UniversitA di Torino, Torino, Italy
**INFM, Dipartimento di Fisica, Politecnico di Torino, Torino, Italy
***Department of Physics, University of Puerto Rico, Mayaguez, PR 00680
ABSTRACT
A master equation formalism is used to model the growth and metastasis of a tumor as a function
of the diffusion and absorption of a nutrient. Healthy and cancerous (C-) cells compete to bind the
nutrient, which is allowed to diffuse starting from a prescribed region. Two thresholds are defined
for the quantity of nutrient bound by the C-cells. If this quantity falls below the lower threshold,
the cell dies, while if it increases above the upper threshold, the cell divides according to a
predefined stochastic mechanism. C-cells migrate when they record a low concentration of free
nutrient in the local environment. The model is formulated in terms of a coupled system of
equations for the cell populations and the free and bound nutrient. This system can be solved by
using the Local Interaction Simulation Approach (LISA), a numerical procedure that permits an
efficient and detailed solution and is easily adaptable to parallel processing. With suitable
parameter variation, the model can describe multiple tumor configurations, ranging from the
classical spheroid with a necrotic core favored by mathematicians to very anisotropic shapes with
inhomogeneous concentrations of the various populations. This is important because the nature of
the anisotropy may be crucial in determining whether and how the cancer metastasizes. The
effects of stochasticity and the presence of additional nutrients or inhibitors can be easily
incorporated.
INTRODUCTION
According to the most recent theories, biological events or even life itself cannot be described by
linear equations, but only as a temporal sequence of discrete events, with the choice at every
bifurcation depending on the local conditions at the time and place. A suitable model for life-
related events like cancer growth must therefore
* include a dependence on the environmental variables and cell population properties;
* introduce stochastic fluctuations of the variables within preordained ranges;
* simulate through a large number of cycles the temporal sequence of life events.
To develop a model as general as possible one needs a deep knowledge of the rules driving cell
life, modelling at first only some very simple processes. As a second step one increases the
complexity of the system by introducing evolution into each parameter, and checking the behavior
of the model after each evolutionary step. This approach is expected to mimic the evolution of the
living world, attempting a correspondence with real life.
According to this methodology, a very basic model for cancer growth consists of two populations
(cancerous and normal cells) competing for a single essential nutrient, represented in the present
case by iron [ 1-3]. In later more realistic treatments, the model will be, of course, generalized to
include additional populations and nutrients, with more complex interaction mechanisms. It is,
however, interesting to observe that even this elementary model succeeds in predicting the basic
features of tumor growth.
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Mat. Res. Soc. Symp. Proc. Vol. 489 ©1998 Materials Research Society