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International Journal of Engineering & Technology, 7 (4.14) (2018) 107-111
International Journal of Engineering & Technology
Website: www.sciencepubco.com/index.php/IJET
Research paper
Callus Formation on Endosperm from Immature and Mature
Fruits of Barringtonia Racemosa
D S M Soder
1
, D N A A Khalid
2
, A Saleh
3
, F Pardi
4
, N J Sidik
5
*
Faculty of Applied Sciences, Universiti Teknologi MARA, 40450 Shah Alam, Selangor, Malaysia
* Corresponding author E-mail: norri536@salam.uitm.edu.my
Abstract
Barringtonia racemosa is mangroves type of plant which had been extensively utilized in conventional practices for relieving ailments of
pain and inflammation. Many studies have been done on ethnobotanical profiles, pharmacological activities and chemical compounds in
Barringtonia racemosa. However, there is a limited study on callogenesis of this plant particularly from different maturity stage of fruits.
The present study is to identify the callogenesis of Barringtonia racemosa from endosperm explants of immature and mature fruits in MS
medium supplemented with different concentrations of hormones 2,4-Dichlorophenoxyacetic acid (2,4-D) (0, 0.5, 1.0, 1.5 and 2.0 mg/L)
and Kinetin (KIN) (0, 0.5, 1.0, 1.5 and 2.0 mg/L). The optimum hormone combination was found in callus grown on endosperm of
immature fruits in MS medium supplemented with 1.5 mg/L 2,4-D and 1.0 mg/L KIN. It was also found that the callus in this treatment
grew profusely with highest fresh weight (0.513 ± 0.022 g), 100% callus induction and friable callus texture. The callus fresh weight on
endosperm explants was higher in immature fruits compared to mature fruits for all the hormone combinations. Therefore, callogenesis
were found more efficient from endosperm explant of immature fruits in Barringtonia racemosa species.
Keywords: Barringtonia racemosa, callogenesis, endosperm, immature fruits, mature fruits
1. Introduction
Medicinal plants have been used extensively by natives or medical
practitioner to treat diseases. Barringtonia racemosa Linn.
(Family Lecythidaceae) is one of the medicinal plants possess
various of bioactivity that engaged to habitat in a wet and watery
area such as by the side of fresh swamps, river banks, lakes, shore
of backwaters and banks of paddy fields. Barringtonia racemosa
fruits have been used to treat a cough, asthma as well as diarrhea
in Sri Lanka, and treat malaria in Africa [1]. Meanwhile, in
Malaysia, the shoots of Barringtonia racemosa is widely
consumed perceiving to gain health benefits. There is also a
practice use a paste of the pounded leaves to reduce the itching
skin [2]. Many studies have been done on different parts of
Barringtonia racemosa towards pharmacological activities such as
antibacterial, anti-tumor, anti-nociceptive, antioxidant, anti-
inflammatory, an alpha-glucosidase inhibitor, anti-fungal, anti-
tuberculosis, anti-arthritic and anti-diarrhoeal [3]. The previous
study also supported that Barringtonia racemosa is a plant-
derived anti-gouty arthritis remedy [4].
On the other hand, the ethnomedicinal uses of the Barringtonia
racemosa fruits are mitigating the pain and inflammation [5] as
well as desirable anti-tumour effect in seed extracts [6]. Locally in
Malaysia, the fruits of Barringtonia racemosa with bitter taste are
eaten fresh as a salad or cooked with turmeric coconut stew. The
immature fruits are easier to be eaten fresh due to soft flesh
meanwhile mature fruits have hard fibrous flesh unpalatable to be
eaten.
The use of in vitro culture technique is important in the
conservation of plant germplasm and in securing valuable
products of medicinal and commercial importance. One of the in
vitro cultures that suitable for mass production is callus culture. In
plant tissue culture, the supplemented nutritional medium with
single or multiple hormones is important in directing the callus
growth [7]. Cytokinins and auxins have extensively been used to
induce callus from plant explants in in vitro medium [8]. The
suitable hormone concentrations used of cytokinin and auxin as
well as source of explant is important in the study of callogenesis
or callus formation.
Nevertheless, there are yet no reports on the comparison of callus
formation from endosperm explants of immature and mature fruits
in Barringtonia racemosa species. Hence, determining the
optimum callus culture is essential in a mass production of cells
that can be used in the further study of somatic embryogenesis and
suspension culture for long-term plant conservation and supply.
2. Methodology
2.1. Plant materials
The endosperm explants source was chosen from 2 type fruits
maturity. Immature and matured fruits of Barringtonia racemosa
were collected at Universiti Kebangsaan Malaysia (UKM), Bangi,
Selangor. The voucher specimen (SK 3191/17) of the sample was
deposited in the Herbarium, Department of Biology, Universiti
Putra Malaysia (UPM), Serdang, Selangor. The immature fruit
selection was chosen from big size ranged 6-8 cm with soft texture
of flesh and pale yellow seeds. The matured fruit selection was
categorized by big size ranged 6-8 cm with hard, fibrous texture of
flesh and pale yellow seeds. The matured fruits were carefully
plucked from the tree before fruits reach over mature (dry fibrous
flesh).