ORIGINAL ARTICLE
Co-occurrence of genes encoding carbapenemase, ESBL,
pAmpC and non-b-Lactam resistance among Klebsiella
pneumonia and E. coli clinical isolates in Tunisia
R. Ben Sallem
1
, B. Laribi
1
, A. Arfaoui
1
, S. Ben Khelifa Melki
2
, H. I. Ouzari
1
, K. Ben Slama
1
, T. Naas
3
and N. Klibi
1
1 Laboratory of Microorganisms and Active Biomolecules, Faculty of Sciences of Tunis, University of Tunis El Manar, Tunis, 2092, Tunisia
2 Service of Biology, Carthagene International Hospital of Tunisia, Tunis, Tunisia
3 Bacteriology-Hygiene Unit, Bic^ etre Hospital, Associated French National Reference Center for Antibiotic Resistance: Carbapenemase-Producing
Enterobacteriaceae, Assistance Publique - H^ opitaux de Paris, Le Kremlin-Bic^ etre, France
Significance and Impact of the Study: Our data revealed multiple co-expressed resistance determinants,
including genes encoding carbapenemases, ESBLs, pAmpCs and non-b-Lactam resistance in all car-
bapenem resistance strains investigated. In the present study, we reported a high prevalence of bla
NDM-1
among K. pneumoniae from Carthagene International Hospital of Tunis, mostly related to
imported cases. These data urge for screening of patients coming abroad and strict contact isolation
measures to stop the further spread in hospital settings.
Keywords
carbapenem, colistin, E. coli, K. pneumoniae,
NDM-1, OXA-48.
Correspondence
Naouel Klibi, Laboratory of Microorganisms
and Active Biomolecules, Faculty of Sciences
of Tunis, University of Tunis El Manar, 2092
Tunis, Tunisia.
E-mail: n_klibi@yahoo.fr
2021/0227: received 11 September 2021,
revised 6 January 2022 and accepted 18
January 2022
doi:10.1111/lam.13658
Abstract
This study aimed to investigate the molecular mechanisms of carbapenem and
colistin resistance in K. pneumoniae and E. coli isolates obtained from hospitalized
patients in Carthagene International Hospital of Tunis. A total of 25
K. pneumoniae and 2 E. coli clinical isolates with reduced susceptibility to
carbapenems were recovered. Susceptibility testing and phenotypic screening tests
were carried out. ESBL, AmpC, carbapenemase and other antibiotic resistance
genes were sought by PCR-sequencing. The presence of plasmid-mediated colistin
resistance genes (mcr-1-8) was examined by PCR and the nucleotide sequence of
the mgrB gene was determined. The analysis of plasmid content was performed by
PCR-Based Replicon Typing (PBRT). The clonality of isolates was assessed by
PFGE and multilocus sequence typing (MLST). All of the isolates produced
carbapenemase activity. They showed a great variation in the distribution of
ESBL, AmpC, carbapenemase and other plasmid-mediated resistance
determinants. K. pneumoniae isolates carried bla
NDM-1
(n = 11), bla
OXA-48
(n = 11), bla
NDM-1
+ bla
OXA-48
(n = 1), bla
NDM-1
+ bla
VIM-1
(n = 1), bla
OXA-204
(n = 1), along with bla
CTX-M
, bla
OXA
, bla
TEM
, bla
CMY
, bla
DHA
and bla
SHV
genes
variants on conjugative plasmid of IncL/M, IncR, IncFII
K
, IncFIB, and IncHI1
types. Three sequence types ST101, ST307 and ST15 were identified. The mgrB
alteration g109a (G37S) was detected in a single colistin-resistant, NDM-1 and
OXA-48-coproducing K. pneumoniae isolate. The two E. coli isolates belonged to
ST95, co-produced NDM-1 and CTX-M-15, and harboured plasmid of IncFII and
IncFIB types. To our knowledge, this is the first report in Tunisia of NDM-1,
OXA-48, and CTX-M-15 coexistence in colistin-resistant K. pneumoniae ST15.
Introduction
The dramatic increase in third-generation cephalosporin-
resistant Enterobacterales correlates with the increased
carbapenem consumption and appears to have driven the
emergence and spread of carbapenem resistance. The
most affected Enterobacterales are Klebsiella pneumoniae
followed by Escherichia coli and the most frequently found
Letters in Applied Microbiology 74, 729--740 © 2022 The Society for Applied Microbiology 729
Letters in Applied Microbiology ISSN 0266-8254
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