Forest Pathology. 2018;48:e12435. wileyonlinelibrary.com/journal/efp
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https://doi.org/10.1111/efp.12435
© 2018 Blackwell Verlag GmbH
1 | INTRODUCTION
Mistletoes are highly specialized perennial, evergreen, flower-
ing semiparasitic angiosperms living on aerial parts of their hosts.
They use root-like structures called haustorium into host tree
branches and trunk (Mathiasen, Nickrent, Shaw, & Watson, 2008)
to take up water and minerals (Glatzel, 1983; Popp & Richter, 1998;
Tennakoon, Chak, Lim, & Bolin, 2014; Watson, 2001; Zuber, 2004),
leading to several physiological and morphological changes in the
tree (Stanton, 2007). The transpiration rate of mistletoe is higher
than that of the host (Mathiasen et al., 2008; Ullmann et al., 1985).
This exacerbates drought stress of trees especially in water-deficit
areas (Glatzel & Geils, 2009), leading to growth loses and acceler-
ated mortality by such secondary factors as fungi and bark beetles
(Hawksworth & Wiens, 1996). Mistletoe can also be responsible for
reduced seed production and low wood quality in the host plants
(Hawksworth & Wiens, 1996; Muir & Hennon, 2007). In addition,
mistletoe infection causes crown degradation by reducing the num-
ber of needles, needle length and width and the longevity of needles
on the infected branches (Rigling, Eilmann, Koechli, & Dobbertin,
Received: 29 December 2017
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Accepted: 15 March 2018
DOI: 10.1111/efp.12435
ORIGINAL ARTICLE
Quantifying the effect of pine mistletoe on the growth of Scots
pine
E. Bilgili
1
| M. Ozturk
1
| K. A. Coskuner
1
| I. Baysal
2
| B. Serdar
1
| H. Yavuz
1
|
M. Eroglu
1
| Y. Usta
1
1
Faculty of Forestry, Karadeniz Technical
University, Trabzon, Turkey
2
Faculty of Forestry, Duzce University,
Duzce, Turkey
Correspondence
Kadir Alperen Coskuner, Faculty of Forestry,
Karadeniz Technical University, Trabzon,
Turkey.
Email: kacoskuner@ktu.edu.tr
Funding information
The Scientific and Technological Research
Council of Turkey (TUBITAK), Grant/Award
Number: 112O258
Editor: H. T. Doğmuş-Lehtijärvi
Summary
Mistletoe infection results in substantial growth losses in mistletoe-infected forests.
This study reports and evaluates the results of retrospective analyses of radial
growth of Scots pine (Pinus sylvestris) in relation to the level of infection of pine mis-
tletoe (Viscum album ssp. austriacum). A total of 43 Scots pine trees were destruc-
tively sampled from different sites. Of these trees, 14 were uninfected and 29 were
infected. Infection classes were determined using six-class dwarf mistletoe rating
system (DMRS). All needle and mistletoe biomass were removed completely and
weighed for each sampled tree. Subsamples from needles and all mistletoe biomass
were taken to the laboratory for oven-dried weight determinations. Five-cm-thick
wood discs were cut from the stem at the breast height (1.3 m) to determine annual
basal area increment for the last 25 years. In addition to DMRS, new infection classes
were created using mistletoe-to-needle biomass (MB/NB) ratio. The results showed
that the radial growth losses could be as much as 41% to 64% at different infection
levels. The rate of growth loss in relation to DMRS and MB/NB ratio was similar, but
with a larger variability in DMRS values. The results showed that both DMRS rating
and MB/NB ratio seem to be important for quantifying growth loss on Scots pine
trees infected with mistletoe. The results of this study can also be invaluable in mod-
elling the effects of mistletoe on the growth of Scots pine trees.
KEYWORDS
Viscum album, Pinus sylvestris, basal area increment, growth loss