Vol.:(0123456789) 1 3
Behavior Genetics
https://doi.org/10.1007/s10519-018-9922-2
ORIGINAL RESEARCH
Efects of Repeated Treatment with Midazolam in SHR and SLA16 Rat
Strains in the Triple Test
Lucía Raily Acuña
1
· Renata Cristina Nunes Marchette
1,2
· Natalli Granzotto
1,2
· Paula Gomes Dias
1
·
Maria Elisa Corvino
1
· Fernando Gabriel Mazur
1
· Fernanda Junques Corrêa
1,2
· Geison Souza Izídio
1
Received: 26 April 2018 / Accepted: 11 September 2018
© Springer Science+Business Media, LLC, part of Springer Nature 2018
Abstract
We exposed male and female rats of SHR (Spontaneously Hypertensive Rats) and SLA16 (SHR.LEW-Anxrr16) strains, in
a non-drugged state, for fve consecutive days to the Triple Test (experiment 1); or after repeated treatment with midazolam
(MDZ), for four consecutive days. The ffth day was performed without treatment (experiment 2). The frst experiment showed
that males did not avoid and females increased the exploration of the open arms over the days. In experiment 2, SLA16 from
both sexes approached more the open arms than SHR rats. The MDZ anxiolytic-like efect was sustained in both strains and
sexes over the days. On the ffth day, SLA16 still approached more the open arms than SHR rats. Data suggest an absence
of repeated-trial tolerance to MDZ anxiolytic-like efects. Testing the SHR and SLA16 strains, especially females, could be
necessary for the future search for the genes and molecular pathways underlying anxiety/emotionality.
Keywords Midazolam · Anxiety · Emotionality · Anxiolytic drugs · Rats
Introduction
There is a need to improve the interpretation of animal and
pre-clinical behavioral tests to evaluate, test and develop new
drugs that will be used in psychiatric disorders treatment.
The elevated plus-maze (EPM) is one of the most com-
monly used animal models of anxiety/emotionality. A naïve
rat or mouse, when exposed to the EPM for the frst time,
will exhibit avoidance of the open arms that is decreased by
classical anxiolytic drugs (i.e., benzodiazepines; BDZs) and
increased by anxiogenic substances (Carobrez and Bertoglio
2005; Handley and Mcblane 1993; Pellow et al. 1985).
However, the response to anxiolytic drugs has been
shown to diminish or disappear when rats are submitted for
one single trial on the EPM. This phenomenon is known
as “One-Trial Tolerance” (OTT) (File 1990). It seems that
rats consolidate some memories related to exploration of
the open arms causing long-term changes in their behavioral
responses. These memories infuence drug responsiveness in
the subsequent trials impairing the search of new anxiolytic
in the preclinical research (for a review see Carobrez and
Bertoglio 2005). The OTT seems to occur independently
of the emotional level of the animals, during the frst expo-
sure to the EPM, the time interval between exposures and
the material of which EPM was built (Cruz-Morales et al.
2002; File 1990; Rodgers and Shepherd 1993). It has been
described not only for benzodiazepines, but also for ethanol,
phenobarbital and NMDA/glycine-B receptor ligands (Rodg-
ers et al. 1992; File and Zangrossi 1993; Treit et al. 1993;
File et al. 1998, 1999; Frussa-Filho et al. 1999; Bertoglio
and Carobrez 2002a, b, 2003).
As commented before, it is well known that the OTT phe-
nomenon involves learning and memory processes and it has
been reported that learning impairment caused by adminis-
tering amnesic doses of BDZs, scopolamine or propanolol
Edited by Stephen Maxson.
Electronic supplementary material The online version of this
article (https://doi.org/10.1007/s10519-018-9922-2) contains
supplementary material, which is available to authorized users.
* Geison Souza Izídio
geisonizidio@gmail.com
1
Laboratório de Genética do Comportamento, Departamento
de Biologia Celular, Embriologia e Genética, Centro
de Ciências Biológicas, Universidade Federal de Santa
Catarina, BEG, bloco C, 3° andar, sala 310C, Florianópolis,
SC CEP 88.040-900, Brazil
2
Department of Pharmacology, Federal University of Santa
Catarina, Florianópolis, Brazil