Vol.:(0123456789) 1 3
Plant Cell Tiss Organ Cult (2017) 130:13–24
DOI 10.1007/s11240-017-1200-0
ORIGINAL ARTICLE
Horseradish esterases: detection, purifcation and identifcation
Ivana Leščić Ašler
1
· Petra Peharec Štefanić
2
· Biljana Balen
2
· Günter Allmaier
3
·
Martina Marchetti‑Deschmann
3
· Biserka Kojić‑Prodić
1
Received: 17 August 2016 / Accepted: 10 March 2017 / Published online: 19 March 2017
© Springer Science+Business Media Dordrecht 2017
Keywords Armoracia lapathifolia · Protein
identifcation · Protein purifcation · Electrophoresis · Mass
spectrometry · Esterase
Introduction
In vitro grown tissues can be used for production of biolog-
ically active compounds. Esterases are known to participate
in many biological processes in plants, but only a handful
of plant esterases are thoroughly characterized. For many
decades now, hydrolases (esterases in particular) have
been studied as valuable tools for stereospecifc hydrolysis
and synthesis of many biotechnologically important com-
pounds. Microbial enzymes have gained interest for their
wide uses in bioconversions related to food, agriculture,
production of fne chemicals, medicine, diagnostics, energy
and waste treatment (Singh et al. 2016; Gurung et al. 2013;
Sayali and Surekha 2013). Esterases have been used in
synthesis of chiral drugs, in producing favoring and fra-
grance compounds, as well as for hydrolysis of milk fat for
the purpose of favor enhancement in the manufacturing of
cheese-related products (Panda and Gowrishankar 2005).
Microbial esterases from baker’s yeast and aroma esters
have been used in production of alcoholic beverages, over
three decades (Suomalainen 1981). The research related to
enzymatic degradation of acetyl-xylan by cooperativity of
fungal esterases and xylanases, published in 1985 (Bieley
et al. 1985), initiated very important studies on decom-
position of biomass which is among the highest priority
in biotechnology these days (Liu and Ding 2016). Bacte-
rial and fungal esterases are generally well-characterized
and widely applied (Borelli and Trono 2015), due to the
simple access to large amounts of these enzymes (since
genetic manipulation with enzymes of microbial origin is
Abstract Our goal is to characterize esterases from
horseradish tissues and assign their physiological roles.
In the present study we focused on isolation, purifcation
and identifcation of esterases from diferent horserad-
ish tissues: plantlets and two tumor tissue lines. Horizon-
tal IEF system enabled separation of six esterase isoforms
with quite diferent pI values as well as with pronounced
diferences in expression levels among analyzed tissues.
Esterases were extracted, fractionated by means of cation
exchange chromatography, and analyzed by planar gel elec-
trophoresis (SDS–PAGE) and isoelectrical focusing (IEF),
UV/Vis spectroscopy, MALDI mass spectrometry (MS)
and MALDI-MS/MS. Several chromatographic strategies
were applied for esterase purifcation and characteriza-
tion. Two subsequent cation exchange chromatographic
steps based on SP-Sepharose FF material, followed by in-
solution digestion combined with MALDI-MS and MS/MS
proved to be the best strategy for identifcation of two ester-
ase proteins, namely Pectinesterase/pectinesterase inhibitor
18 and GDSL esterase/lipase ESM1.
* Ivana Leščić Ašler
ilescic@irb.hr
1
Department of Physical Chemistry, Laboratory for Chemical
and Biological Crystallography, Ruđer Bošković Institute,
Bijenička cesta 54, 10000 Zagreb, Croatia
2
Division of Molecular Biology, Department of Biology,
Faculty of Science, University of Zagreb, Horvatovac 102a,
10000 Zagreb, Croatia
3
Institute for Chemical Technologies and Analytics,
Technische Universität Wien (Vienna University
of Technology), Getreidemarkt 9/164, 1060 Vienna, Austria