Vol.:(0123456789) 1 3
Environmental Sustainability
https://doi.org/10.1007/s42398-020-00146-6
ORIGINAL ARTICLE
Response of a rice‑feld cyanobacterium Anabaena sp. HKAR‑7
upon exposure to ultraviolet‑B radiation and ammonium chloride
Deepak Kumar Singh
1,2
· Jainendra Pathak
3
· Abha Pandey
1
· Vidya Singh
1
· Haseen Ahmed
1,4
·
Deepak Kumar
1
· Rajneesh
1
· Rajeshwar Prasad Sinha
1
Received: 26 November 2019 / Revised: 30 August 2020 / Accepted: 3 November 2020
© Society for Environmental Sustainability 2020
Abstract
Interactive efects of ultraviolet radiation (UVR), photosynthetically active radiation (PAR) and exogenously supplied ammo-
nium chloride (NH
4
Cl) was studied in the rice-feld cyanobacterium Anabaena sp. HKAR-7. The cyanobacterium was cul-
tured under varying NH
4
Cl concentrations i.e., 0, 50, 200, 500, 1000 and 5000 μM and 200 μM (concentration) was found to
be optimum for the growth of the cyanobacterium. Detrimental efects of UV-B exposure were observed on photosynthetic
pigments such as chlorophyll a (Chl a), carotenoids and phycocyanin (PC). However, damage to these pigments was less
in the cyanobacterial samples supplemented with NH
4
Cl. Contents of Chl a and PC in cyanobacterial cells decreased upon
UV-B exposure but decrement was less in the samples supplemented with NH
4
Cl. Upon UV-B exposure, carotenoids content
enhanced initially (till 15 days) during the course of treatment (21 days) but signifcant decrease (in carotenoids content) was
observed in later phase of the experiment. From the results of photosynthetic activity, maximum quantum efciency of PSII
(Fv/Fm) and maximum electron transport rate (ETR
max
), it could be concluded that exogenous supplementation of NH
4
Cl
(optimum concentration) helped in protecting the cyanobacterial cells from highly energetic UVR to certain extent. Another
interesting observation was signifcantly higher levels of biosynthesis and accumulation of mycosporine-like amino acids
(MAAs) in the cyanobacterial cells supplemented with NH
4
Cl in comparison to non-supplemented cells. The purifed MAA
was identifed to be phorphyra-334 as evidenced by UV/VIS absorption spectra, high performance liquid chromatography
(HPLC) and electrospray ionization-mass spectrometry (ESI–MS).
Keywords Ammonium chloride · Cyanobacteria Anabaena · Mycosporine-like amino acids · Photoprotection · Ultraviolet
radiation
Introduction
Ultraviolet radiation (UVR), the comparatively low wave-
band radiation, is composed of highly energetic photons
which reach the Earth’s surface along with solar radiation.
In current scenario, UVR infux has increased on the Earth
due to anthropogenically released ozone depleting com-
pounds (Häder et al. 2015). Although, UV-B constitute
less than 1% of the total incoming solar radiation (Vincent
and Roy 1993), it severely afects crucial biomolecules
such as DNA, RNA and proteins which are important for
biochemical, physiological and genetic functioning of
the cell (Sinha and Häder 2016; Rajneesh et al. 2019).
Besides, in cyanobacteria, detrimental efects of UVR on
pigmentation, phycobiliprotein composition, motility, N
2
metabolism, DNA, protein profle and
14
CO
2
uptake have
been well documented (Kannaujiya and Sinha 2015; Sinha
Electronic supplementary material The online version of this
article (https://doi.org/10.1007/s42398-020-00146-6) contains
supplementary material, which is available to authorized users.
* Rajeshwar Prasad Sinha
r.p.sinha@gmx.net; rpsinhabhu@gmail.com
1
Laboratory of Photobiology and Molecular Microbiology,
Centre of Advanced Study in Botany, Institute of Science,
Banaras Hindu University, Varanasi 221005, India
2
Department of Botany, Acharya Narendra Deo Kisan
P. G. College, Gonda 271313, India
3
Department of Botany, Pt. Jawaharlal Nehru College,
Banda 210001, India
4
Department of Botany, Government Girls PG College,
Satna 485001, India