Research Article
Investigation of Ser315 Substitutions within
katG Gene in Isoniazid-Resistant Clinical Isolates of
Mycobacterium tuberculosis from South India
A. Nusrath Unissa, N. Selvakumar, Sujatha Narayanan, C. Suganthi, and L. E. Hanna
Division of Biomedical Informatics, Department of Clinical Research, National Institute for Research in Tuberculosis (NIRT),
Indian Council of Medical Research (ICMR), No. 1 Mayor Sathyamoorthy Road, Chetput, Chennai, Tamil Nadu 600 031, India
Correspondence should be addressed to L. E. Hanna; hanna@trcchennai.in
Received 2 July 2014; Accepted 20 October 2014
Academic Editor: Filippo Canducci
Copyright © 2015 A. Nusrath Unissa et al. Tis is an open access article distributed under the Creative Commons Attribution
License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly
cited.
Mutation at codon 315 of katG gene is the major cause for isoniazid (INH) resistance in Mycobacterium tuberculosis (M. tuberculosis).
Substitution at codon 315 of katG gene was analyzed in 85 phenotypically resistant isolates collected from various parts of southern
India by direct sequencing method. Te obtained results were interpreted in the context of minimum inhibitory concentration
(MIC) of INH. Of the 85 phenotypically resistant isolates, 56 (66%) were also correlated by the presence of resistance mutations in
the katG gene; 47 of these isolates had ACC, 6 had AAC, 2 had ATC, and one had CGC codon. Te frequency of Ser315 substitution
in katG gene was found to be higher (70%) amongst multidrug-resistant (MDR) strains than among non-MDR (61%) INH-resistant
isolates. Further, the frequency of mutations was found to be greater (74%) in isolates with higher MIC values in contrast to those
isolates with low MIC values (58%). Terefore, the study identifed high prevalence of Ser315Tr substitution in katG gene of INH-
resistant isolates from south India. Also, isolates harboring this substitution were found to be associated with multidrug and high
level INH resistance.
1. Introduction
Although tuberculosis (TB) is a preventable, treatable, and
curable disease, it still remains as a major public health
problem. Te period of 6–9 months needed to treat the
disease is too long, and the treatment is ofen associated with
signifcant toxicity. Tese factors make patient compliance
to therapy very difcult, leading to the emergence and
selection of drug-resistant TB bacteria [1]. Te emergence
of multidrug-resistant (MDR) TB, defned as strains which
are resistant to two most potent anti-TB drugs, namely,
isoniazid (INH) and rifampicin (RIF), and XDR-TB, defned
as MDR-TB strains that are resistant to second-line TB drugs,
that is, fuoroquinolones and at least one of the injectable
aminoglycosides (capreomycin, kanamycin) or amikacin,
has worsened the situation further [2]. Globally, 450,000
people developed MDR-TB in 2012; more than half of these
cases were from India, China, and the Russian Federation.
It is estimated that about 9.6% of MDR-TB cases had XDR-
TB. About 170,000 MDR-TB deaths are estimated to have
occurred in 2012 [3].
Resistance against all known anti-TB drugs has been
reported. However, isolates of M. tuberculosis resistant to INH
are seen with increasing frequency (1 in 10
6
) as compared
to isolates resistant to other drugs [4]. Also, resistance to
INH, alone or in combination with other drugs, is now the
second most common cause for resistance. Globally, it has
been estimated that INH resistance was found in 10.3% of new
cases and 27.7% of treated cases [5]. An earlier study based
on susceptibility testing from south India also reported high
resistance to INH in 15.4% of cases compared to 4.4% cases
of RIF resistance [6].
INH has been used extensively as the frontline anti-TB
drug and a drug of choice for chemoprophylaxis, acting as
a principle component in the current six-month short course
chemotherapy regimen. It has long been recognized that INH
Hindawi Publishing Corporation
BioMed Research International
Volume 2015, Article ID 257983, 5 pages
http://dx.doi.org/10.1155/2015/257983