J Fish Dis. 2019;00:1–11. wileyonlinelibrary.com/journal/jfd
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1 © 2019 John Wiley & Sons Ltd
1 | INTRODUCTION
Chile is the second largest producer of farmed salmon and trout
in the world after Norway (FAO, 2011). The national and interna‐
tional salmon farming industry is mainly threatened by extensive
losses caused by infectious and contagious diseases occurring
during the production cycle. Salmonid rickettsial septicaemia (SRS)
or Piscirickettsiosis is one of the leading diseases affecting salmon
reared in net pens in Chile, whose bacterial causative agent is
Piscirickettsia salmonis, a Gram‐negative, obligate, intracellular,
non‐motile and non‐encapsulated pathogen characterized by a
variable diameter ranging between 0.5 and 1.5 µm. Piscirickettsia
salmonis isolate LF‐89, described by Fryer, Lannan, Giovannoni, and
Wood (1992), is classified as a reference strain (Cvitanich, Garate
n., & Smith, 1991; Fryer, Lannan, Garcés, Larenas, & Smith, 1990).
Piscirickettsiosis affects different varieties of salmon species, mainly
Atlantic salmon (Salmon salar L.), rainbow trout (Oncorhynchus mykiss
W.) and Coho salmon (Oncorhynchus kisutch W.).
Antibiotics have been used for many years as the first tool in
the prevention and treatment of this infectious and contagious
disease, being florfenicol and oxytetracycline by far the two most
commonly used antimicrobials in Chile (Grave, Lingaas, Bangen,
Received: 3 June 2019
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Revised: 2 August 2019
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Accepted: 5 August 2019
DOI: 10.1111/jfd.13089
ORIGINAL ARTICLE
Analysis of single nucleotide polymorphisms (SNPs) associated
with antibiotic resistance genes in Chilean Piscirickettsia
salmonis strains
Jaime Figueroa
1,2
| Diana Castro
2
| Fernando Lagos
2
| Carlos Cartes
2
|
Adolfo Isla
2
| Alejandro J. Yáñez
1,2
| Rubén Avendaño‐Herrera
2,3
| Denise Haussmann
4
1
Faculty of Sciences, Institute of
Biochemistry and Microbiology, Universidad
Austral de Chile, Valdivia, Chile
2
FONDAP Centre, Interdisciplinary
Centre for Aquaculture Research (INCAR),
Concepción, Chile
3
Laboratory of Pathology of
Aquatic Organisms and Aquaculture
Biotechnology, Faculty of Life
Sciences, Universidad Andrés Bello, Viña del
Mar, Chile
4
Department of Basic Sciences, Faculty of
Sciences, Universidad Santo Tomás, Valdivia,
Chile
Correspondence
Jaime Figueroa, Faculty of Sciences,
Institute of Biochemistry and Microbiology,
Universidad Austral de Chile, Valdivia, Chile.
Email: jefigueroa@uach.cl
Funding information
Interdisciplinary Centre for Aquaculture
Research (INCAR) at Universidad Austral de
Chile, Grant/Award Number: 122929 and
15110027
Abstract
The aetiological agent of Piscirickettsiosis is Piscirickettsia salmonis, a Gram‐negative
intracellular pathogen, and high doses of antibiotics have regularly been employed
to treat this infection. Seven florfenicol and/or oxytetracycline resistance genes (tet
pump, tetE, Tclor/flor , Tbcr , TfloR, ompF and mdtN) were identified in strains by in silico
genome analyses. Later, the number of single nucleotide polymorphisms (SNPs) and
its relationship with the resistance to these antibiotics were identified and analysed,
using the original LF‐89 strain as reference. Trials to determine and compare the
minimum inhibitory concentration (MIC) of oxytetracycline and florfenicol in each
strain, as well as to quantify the gPCR transcripts levels in the selected genes, were
performed. Therefore, variations in the resistance to both antibiotics were observed,
where the strain with fewer SNPs showed the highest susceptibility. Consistently,
the in silico 3D analyses of proteins encoded by the selected genes revealed struc‐
tural changes, evident in the sequences with the highest number of SNPs. These re‐
sults showed that the bacterial resistance to oxytetracycline was mainly linked to the
presence of SNPs in relevant sites, antibiotic resistance genes and an OmpF porin,
leading to important changes in the protein structure.
KEYWORDS
florfenicol, oxytetracycline, Piscirickettsiosis, resistance genes, SNPs