SEROLOGIC EVIDENCE OF INFECTION WITH EHRLICHIAE AND SPOTTED
FEVER GROUP RICKETTSIAE AMONG RESIDENTS OF GAG ISLAND, INDONESIA
ALLEN L. RICHARDS, SUTANTI RATIWAYANTO, EKO RAHARDJO, DARYL J. KELLY, GREGORY A. DASCH,
DAVID J. FRYAUFF, AND MICHAEL J. BANGS
U.S. Naval Medical Research Unit No. 2, Jakarta, Indonesia; Communicable Diseases Research Center, National Institute of Health
Research and Development, Jakarta, Indonesia; U.S. Naval Medical Research Center, Silver Spring, Maryland; Walter Reed Army
Institute of Research, Washington,DistrictofColumbia
Abstract. The causative agents of scrub and murine typhus are considered endemic to Indonesia. However, the
presence of spotted fever group rickettsiae and ehrlichiae have not been previously described in this country. During an
investigation of arthropod-borne diseases on Gag Island, located northwest of the island of New Guinea in eastern
Indonesia, the prevalence of antibody to the etiologic agents of monocytic ehrlichiosis, spotted fever rickettsiosis, and
scrub and murine typhus were determined. Analysis of 55 blood samples from residents of Gag Island showed serore-
activity to antigen preparations of Ehrlichia chaffeensis (7 of 48, 14.6%), two spotted fever group rickettsiae: Rickettsia
rickettsii (5 of 48, 10.4%) and R.conorii (10 of 49, 20.4%), Orientiatsutsugamushi (5 of 53, 9.4%), and R.typhi (1 of 48,
2.1% [by an indirect immunofluorescence assay] and 1 of 50, 2.0% [by an enzyme-linked immunosorbent assay]). These
results show serologic evidence of infection with ehrlichiae and spotted fever group rickettsiae for the first time in
Indonesia in a location where the prevalence of antibody to O. tsutsugamushi and R. typhi was lower.
INTRODUCTION
Rickettsial diseases are endemic to Indonesia.
1
Murine ty-
phus (Rickettsia typhi) has been reported on many of the
larger islands of the archipelago, including Java,
2–7
Sumatra,
8,9
Bali,
4,10
Muna,
4
and Irian Jaya.
11
The closely re-
lated etiologic agent of epidemic typhus, R. prowazekii, does
not occur in Indonesia
2,4,12,13
due to the absence of the ar-
thropod vector, the human body louse Pediculus humanus
humanus.
2
Scrub typhus (Orientia tsutsugamushi) has been
recognized throughout Indonesia, especially during World
War II when the occurrence of the disease on the island of
New Guinea and neighboring islands was found to be a major
impediment for military forces.
14–16
In addition to New
Guinea, scrub typhus has been reported on the islands of
Java,
5–7,17–19
Sumatra,
8,9
Borneo,
20
and Sulawesi.
21
Other
rickettsiae and Rickettsia-like organisms, such as spotted fe-
ver group rickettsiae and ehrlichiae, have not been previously
reported occurring in Indonesia,
1,13,22
although they have
been reported in the surrounding Asia-Pacific region, includ-
ing Australia,
12,23–25
Malaysia,
22,26–29
Thailand,
30–34
Ja-
pan,
32,35–38
India,
12,31
Pakistan,
30,31
Nepal,
31
China,
39,40
Mon-
golia,
31
and Taiwan.
32
This report was the outcome of an investigation to deter-
mine the presence and level of arthropod-borne diseases on
Gag Island, a small, isolated island located northwest of the
Province of Irian Jaya, Indonesia. An investigation on malaria
prevalence and parasite drug resistance has been reported
elsewhere.
41
Herein we describe, for the first time, serologic
evidence of life-long inhabitants of Indonesia infected with
spotted fever group rickettsiae and ehrlichiae. Additionally,
we report a lower prevalence of antibody to O.tsutsugamushi
and a near absence of seroreactivity to R. typhi among resi-
dents of Gag Island, Indonesia.
MATERIALS AND METHODS
Study population. Gag is a small island located in the Hal-
mahera Sea, northwest of Irian Jaya and along the eastern
edge of the Maluku Island Chain in the Republic of Indonesia
(Figure 1). Approximately, 500 indigenous people lived on
this island at the time of the survey, most of them in the
village of Gambir. In addition, approximately 200 non-
indigenous mining personnel were temporarily residing on
the island. The local inhabitants were primarily subsistence
farmers and fishermen. Gambir is located on the coast along
the northeast side of the island, surrounded by sago swamp,
coconut groves, and native forest. Most people lived in well-
constructed, cement homes; however, the houses still allowed
easy excess to rodents and arthropod vectors. Four hundred
fifty-six residents of Gambir and 112 mining employees were
screened for malaria and lymphatic filariasis. Of these, 80
villagers and three mining employees were enrolled in a study
of in vivo resistance of malaria parasites to chloroquine and
Fansidar® (F. Hoffmann-La Roche, Basel, Switzerland).
41
From these 83 individuals, 55 sera (53 from villagers and two
from mining employees) were available for testing of expo-
sure to rickettsiae and rickettsia-like organisms.
Human blood was collected by venipuncture, separated,
and the serum was stored at -70°C. For the indirect immuno-
fluorescence assay (IFA), the serum was assayed for reactiv-
ity to R. typhi, R. rickettsii, and E. chaffeensis as describe by
the manufacturer (MRL Diagnostics, Cypress, CA). Sera
were also assessed for reactivity to R. typhi, R.conorii, and O.
tsutsugamushi with an enzyme-linked immunosorbent assay
(ELISA) format previously described.
6
This study was reviewed and approved by the Committee
for the Protection of Human Subjects of the U.S. Naval Medi-
cal Research Unit No. 2 and the Communicable Disease Re-
search Center, National Institutes of Health, Research and
Development (Jakarta, Indonesia). Informed consent was ob-
tained from all human adult participants and from parents or
legal guardians of minors.
RESULTS
Fifty-five human serum specimens from Gag Island were
examined for serologic reactivity to E. chaffeensis, R. rick-
ettsii,R.conorii,O.tsutsugamushi, and R.typhi antigen prepa-
rations. The seroreactivity prevalence is shown in Table 1.
Agreement was found between the results obtained with dif-
ferent assays (IFA and ELISA) for detecting antibodies to
Am. J. Trop. Med. Hyg., 68(4), 2003, pp. 480–484
Copyright © 2003 by The American Society of Tropical Medicine and Hygiene
480